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Montevideo spike rush, sand spikerush

ambiguous spikesedge, éléocharide ambiquë

Habit Plants perennial, mat-forming; rhizomes evident, long, 0.7–2 mm thick, firm, cortex persistent, longer internodes 1–2 cm, scales often fugaceous, 6–8 mm, membranous, not fibrous. Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 1–3 cm, scales persistent, 5–10 mm, membranous, not fibrous.
Culms

terete or cross section elliptic or rectangular, with 5–10 blunt ridges when dry, 25–50 cm × 0.5–1.2 mm, firm to hard, spongy.

terete, often with to 16 blunt ridges when dry, 25–80 cm × 0.5–1.5 mm, firm, internally spongy.

Leaves

distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery, apex usually red-brown, often callose, subtruncate to obtuse, tooth usually present on some or all culms, to 0.9 mm.

distal leaf sheaths persistent, not splitting, proximally dark red, distally red to stramineous or green, often callose, thinly papery (to membranous), apex dark red-brown, obtuse to subtruncate, tooth sometimes present on some culms, to 0.3 mm.

Spikelets

usually ovoid or ellipsoid to subcylindric, rarely lanceoloid, 4–12 × (1.5–)2–3 mm, apex rounded to acute;

proximal scale amplexicaulous or clasping over 3/4 of culm, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 30–100, 6–10 per mm of rachilla, orange-brown, midrib regions often greenish, oblong to ovate, 1.5–2.5 × 1–1.5 mm, apex entire, broadly rounded, sometimes acute in distal part of spikelet, usually horizontally wrinkled and recurved, often carinate in distal part of spikelet.

ovoid to lanceoloid, 5–23 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 10–60, 3–4 per mm of rachilla, medium brown, rarely red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, in distal part lanceolate, 2.5–3.5 × 1.7 mm, entire, apex acute to rarely obtuse, often carinate in distal part of spikelet.

Flowers

perianth bristles 5–6(–7), stramineous to medium brown, stout, often unequal, much shorter than to equaling achene, rarely all rudimentary;

stamens 3;

anthers dark yellow to stramineous, 0.8–1.5 mm;

styles 3-fid or some 2-fid.

perianth bristles 2–4(–5), brown, slender to stout, usually very unequal, rudimentary to equaling achene;

stamens 3;

anthers brown, 1.5–2 mm;

styles 2-fid, sometimes some 3-fid.

Achenes

falling with scales, dark brown, obovoid to obpyriform, compressed trigonous, angles evident to obscure, 0.7–1 × 0.65–0.8 mm, neck absent to sometimes long, finely rugulose at 10–30X, over 20 horizontal ridges in vertical series, and/or minutely cancellate at 20–30X.

not persistent, dark yellow or stramineous, obovoid to obpyriform, all biconvex or some (very rarely all) compressedtrigonous, angles obscure, 1–1.5 × 0.9–1.25 mm, apex rounded, neck absent (to very short), finely rugulose at 10–20X, with 20 or more horizontal ridges in vertical series and/or finely cancellate at 10–30X.

Tubercles

brown to whitish, pyramidal, as high as wide or sometimes greatly depressed, (0.1–)0.25–0.3 × 0.2–0.3 mm.

brown or proximally whitish, pyramidal, depressed, rarely some as high as wide, 0.15–0.3(–0.5) × (0.3–)0.4–0.6 mm.

2n

= 20.

= 44, 45, 46.

Eleocharis montevidensis

Eleocharis ambigens

Phenology Fruiting spring–fall. Fruiting spring–summer.
Habitat Wet soil, fresh ponds, lakes, streams, springs, seeps, marshes, ditches, grasslands Coastal (rarely inland) fresh to brackish pond shores, marshes, disturbed places
Elevation 0–2300 m (0–7500 ft) 0–10 m (0–0 ft)
Distribution
from FNA
AL; AZ; CA; FL; KS; LA; MS; NC; NM; OK; SC; TX; Mexico; South America
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; FL; GA; LA; MA; MS; NC; NJ; RI; SC; TX; VA; QC
Discussion

The name Eleocharis montana was long misapplied to E. montevidensis. The type of E. palmeri, recognized as a species (H. K. Svenson 1957), is indistinguishable from specimens of E. montevidensis. Eleocharis montevidensis is extremely variable; the broadly rounded, usually wrinkled and recurved floral scales are diagnostic. Achenes often fail to form. Eleocharis montevidensis is apparently closely related to E. parishii, in which the floral scales are less densely placed on the rachilla and acute to rounded, and the spikelets are narrowly lanceoloid to cylindric. In the absence of achenes and rhizomes, some specimens of E. montevidensis are easily mistaken for E. tricostata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis ambigens is usually treated as a synonym of E. fallax because some specimens from Massachusetts to Georgia have some 3-fid styles and some obscurely trigonous achenes. A specimen of E. ambigens from Louisiana has trigonous achenes. Eleocharis ambigens is very similar to and perhaps better included in E. uniglumis (S.-O. Strandhede 1967). Intermediates with E. macrostachya occur in Louisiana and Texas. The depressed tubercles are diagnostic of E. ambigens. Eleocharis ambigens is sometimes confused with E. montevidensis, which is readily distinguished by its floral scales with broadly rounded apices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 80. FNA vol. 23, p. 78.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. arenicola, E. montana subsp. montevidensis, E. palmeri
Name authority Kunth: Enum. Pl. 2: 144. (1837) Fernald: Rhodora 37: 394. (1935)
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