Eleocharis microcarpa |
Eleocharis torticulmis |
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small-fruit spikesedge, smallfruit spikerush |
California twisted spikerush, twist-stem spikerush |
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Habit | Plants usually annual, tufted, sometimes mat-forming via proliferating and arching inflorescences (stoloniferous), sometimes entirely vegetative; rhizomes absent. | Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick. | ||||
Culms | often ascending or arching, quadrangular or broadly elliptic, 2–40 cm × 0.1–0.4(–0.6) mm, soft. |
erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm; culm tufts not proximally bulbous. |
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Leaves | distal leaf sheaths persistent or disintegrating, pale brown, green or red-brown, streaked or mottled red-brown or purple, translucent, membranous, apex narrowly acute. |
distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse. |
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Spikelets | basal spikelets absent; often proliferous, ovoid to ellipsoid or lanceoloid, terete, 2–10.7 × 1–2 mm, apex acute; proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, often longer, often resembling an involucral bract, ovate to lanceolate, midrib markedly thickened and broad and often prolonged beyond lamina; subproximal scale with flower; floral scales spiraled, colorless, whitish, or pale brown, streaked or mottled red-brown or purple, midribs green and red-brown or green and purple, ovate to elliptic, 0.8–1.5 × 0.4–0.8 mm, membranous, midrib obscure to prominent. |
6–8 × 2–3 mm; proximal scale empty, 3–4 mm, shorter than to equaling spikelet; floral scales 8–10 per spikelet, 3.5–5 × 2 mm. |
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Flowers | perianth bristles present or sometimes apparently absent; stamens 3; anthers 0.15–0.35; styles 3-fid. |
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Perianth | bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose. |
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Achenes | whitish to olive or pale brown, sometimes spotted olive or red-brown, obovoid, trigonous (or subterete), angles prominent, 0.55–0.8 × 0.3–0.5 mm, apex constricted proximal to tubercle, smooth. |
stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm; beak 0.3–0.6 mm. |
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Tubercles | green or pale brown to red-brown, trigonous. |
0.25–0.6 × 0.3–0.55 mm. |
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Anthers | 1.8 –3 mm. |
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Eleocharis microcarpa |
Eleocharis torticulmis |
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Phenology | Fruiting summer (Jun–Jul). | |||||
Habitat | Fens, wet meadows, vernal ponds | |||||
Elevation | 1100 m (3600 ft) | |||||
Distribution |
AL; FL; GA; IN; LA; MA; MD; MI; MS; NC; NJ; PA; SC; TN; TX; VA; West Indies
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CA |
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Discussion | Varieties 2 (2 in the flora). Specimens of Eleocharis brittonii and E. microcarpa without achenes are often difficult to distinguish. More detailed study of the four apparent entities across the two species is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 95. | FNA vol. 23, p. 115. | ||||
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis > subg. Zinserlingia | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York 3: 312. (1836) | S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001) | ||||
Web links |