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small-fruit spikesedge, smallfruit spikerush

Suksdorf spikerush, Suksdorf's spike-rush

Habit Plants usually annual, tufted, sometimes mat-forming via proliferating and arching inflorescences (stoloniferous), sometimes entirely vegetative; rhizomes absent. Plants perennial; rhizomes 0.5–1.5 mm thick, scales persistent, 5–10 mm, thinly membranous, not fibrous; resting buds absent (non-resting buds on rhizome apex ellipsoid, 10 × 2–5 mm); caudices present, hard, 2 mm thick.
Culms

often ascending or arching, quadrangular or broadly elliptic, 2–40 cm × 0.1–0.4(–0.6) mm, soft.

erect, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times as wide as thick, (5–)10–40 cm × 0.5–1.2 mm, firm to hard, finely many-ridged at 10–20X;

culm tufts not proximally bulbous.

Leaves

distal leaf sheaths persistent or disintegrating, pale brown, green or red-brown, streaked or mottled red-brown or purple, translucent, membranous, apex narrowly acute.

distal leaf sheaths stramineous to brown or reddish proximally, green to stramineous distally, subtruncate to obtuse, membranous to papery;

apex brown to red.

Spikelets

basal spikelets absent; often proliferous, ovoid to ellipsoid or lanceoloid, terete, 2–10.7 × 1–2 mm, apex acute;

proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, often longer, often resembling an involucral bract, ovate to lanceolate, midrib markedly thickened and broad and often prolonged beyond lamina;

subproximal scale with flower;

floral scales spiraled, colorless, whitish, or pale brown, streaked or mottled red-brown or purple, midribs green and red-brown or green and purple, ovate to elliptic, 0.8–1.5 × 0.4–0.8 mm, membranous, midrib obscure to prominent.

5–10 × 2–4 mm;

proximal scale usually empty, 3–5 mm, usually 1/2 or more as long as spikelet;

floral scales 8–12 per spikelet, lanceolate (to ovate), 3.5–5 × 2–2.5 mm.

Flowers

perianth bristles present or sometimes apparently absent;

stamens 3;

anthers 0.15–0.35;

styles 3-fid.

perianth bristles 6, equal, the longest equaling achene to exceeding tubercle, very slender, spinules dense;

anthers 1.6–3.5 mm.

Achenes

whitish to olive or pale brown, sometimes spotted olive or red-brown, obovoid, trigonous (or subterete), angles prominent, 0.55–0.8 × 0.3–0.5 mm, apex constricted proximal to tubercle, smooth.

medium brown to gray or dark brown, equilaterally trigonous to compressed trigonous, rarely some biconvex, 2–2.7 × 0.7–1.3 mm, apex tapered to a distinct stramineous beak 0.2–0.9 × 0.3–0.6 mm.

Tubercles

green or pale brown to red-brown, trigonous.

0.4–0.5 × 0.3–0.5 mm.

Eleocharis microcarpa

Eleocharis suksdorfiana

Phenology Fruiting late spring–summer.
Habitat Bogs, fens, wet meadows, springs, wet gravel near ponds
Elevation (0–)1100–3300 m ((0–)3600–10800 ft)
Distribution
from FNA
AL; FL; GA; IN; LA; MA; MD; MI; MS; NC; NJ; PA; SC; TN; TX; VA; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; AB; BC
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Specimens of Eleocharis brittonii and E. microcarpa without achenes are often difficult to distinguish. More detailed study of the four apparent entities across the two species is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Eleocharis suksdorfiana is usually included in E. quinqueflora, it clearly differs qualitatively as given in the key. A collection from hot springs in Ruby Valley, Elko County, Nevada, has stout perianth bristles less than half of the achene length and may represent an undescribed taxon related to E. suksdorfiana. Eleocharis suksdorfiana closely resembles E. rostellata in its achenes, tubercles, culms, and caudices; it differs in the presence of long horizontal rhizomes and the absence of stoloniferous culms. The achenes of E. suksdorfiana are often finely longitudinally ridged, but in E. rostellata they are often rugulose. Specimens from Coconino and Santa Cruz counties, Arizona, are probably E. suksdorfiana but lack achenes so cannot be identified with certainty.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianth bristles 0.2–0.4 mm, shorter than achene; proximal scale of spikelets 0.4–0.5 mm wide.
var. microcarpa
1. Perianth bristles 0.6–1 mm, shorter than to equaling achene; proximal scale of spikelets 0.8–1 mm wide.
var. filiculmis
Source FNA vol. 23, p. 95. FNA vol. 23, p. 115.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Subordinate taxa
E. microcarpa var. filiculmis, E. microcarpa var. microcarpa
Synonyms E. paucifora var. suksdorfiana, E. quinqueflora var. suksdorfiana
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 312. (1836) Beauverd: Bull. Soc. Bot. Genève 13: 267. (1922)
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