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small-fruit spikesedge, smallfruit spikerush

daggerleaf spikerush

Habit Plants usually annual, tufted, sometimes mat-forming via proliferating and arching inflorescences (stoloniferous), sometimes entirely vegetative; rhizomes absent.
Culms

often ascending or arching, quadrangular or broadly elliptic, 2–40 cm × 0.1–0.4(–0.6) mm, soft.

20–35 cm × 0.3–1 mm.

Leaves

distal leaf sheaths persistent or disintegrating, pale brown, green or red-brown, streaked or mottled red-brown or purple, translucent, membranous, apex narrowly acute.

apex of distal leaf sheath subacute to narrowly acute, tooth to 0.3(–1.1) mm.

Spikelets

basal spikelets absent; often proliferous, ovoid to ellipsoid or lanceoloid, terete, 2–10.7 × 1–2 mm, apex acute;

proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, often longer, often resembling an involucral bract, ovate to lanceolate, midrib markedly thickened and broad and often prolonged beyond lamina;

subproximal scale with flower;

floral scales spiraled, colorless, whitish, or pale brown, streaked or mottled red-brown or purple, midribs green and red-brown or green and purple, ovate to elliptic, 0.8–1.5 × 0.4–0.8 mm, membranous, midrib obscure to prominent.

narrowly lanceoloid, 3–12 × (1–)2–2.5(–4) mm, apex acute;

floral scales 25–100, 10–12 per mm of rachilla, orangebrown to stramineous, ovate, 1.5–2 × 1 mm, midribs mostly keeled, apex acute or narrowly rounded in proximal part of spikelet.

Flowers

perianth bristles present or sometimes apparently absent;

stamens 3;

anthers 0.15–0.35;

styles 3-fid.

perianth bristles 6–7, brown, stout, the longest equaling or exceeding tubercle;

stamens 2–3;

anthers brown, 0.3 mm;

styles 2-fid.

Achenes

whitish to olive or pale brown, sometimes spotted olive or red-brown, obovoid, trigonous (or subterete), angles prominent, 0.55–0.8 × 0.3–0.5 mm, apex constricted proximal to tubercle, smooth.

0.9–1.1 × 0.6–0.8 mm.

Tubercles

green or pale brown to red-brown, trigonous.

deltoid, 0.25–0.5 × 0.5–0.7 mm, 1/2–2/3 as high as wide, 1/4–1/2 as high and 2/3–4/5 as wide as achene.

2n

= 10.

Eleocharis microcarpa

Eleocharis lanceolata

Phenology Fruiting summer to fall.
Habitat Fresh shores, stream beds, pine woods, disturbed places
Elevation 0–400 m (0–1300 ft)
Distribution
from FNA
AL; FL; GA; IN; LA; MA; MD; MI; MS; NC; NJ; PA; SC; TN; TX; VA; West Indies
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CA; KS; LA; MO; OK; TN; TX
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Specimens of Eleocharis brittonii and E. microcarpa without achenes are often difficult to distinguish. More detailed study of the four apparent entities across the two species is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some collections of Eleocharis lanceolata from Tennessee appear to be intermediate with E. obtusa. The California record is an introduced rice-field weed collected in 1949.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perianth bristles 0.2–0.4 mm, shorter than achene; proximal scale of spikelets 0.4–0.5 mm wide.
var. microcarpa
1. Perianth bristles 0.6–1 mm, shorter than to equaling achene; proximal scale of spikelets 0.8–1 mm wide.
var. filiculmis
Source FNA vol. 23, p. 95. FNA vol. 23, p. 104.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Subordinate taxa
E. microcarpa var. filiculmis, E. microcarpa var. microcarpa
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 312. (1836) Fernald: Proc. Amer. Acad. Arts 34: 493, figs. 27–29. (1899)
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