Eleocharis microcarpa |
Eleocharis bolanderi |
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small-fruit spikesedge, smallfruit spikerush |
Bolander's spikerush |
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Habit | Plants usually annual, tufted, sometimes mat-forming via proliferating and arching inflorescences (stoloniferous), sometimes entirely vegetative; rhizomes absent. | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | ||||
Culms | often ascending or arching, quadrangular or broadly elliptic, 2–40 cm × 0.1–0.4(–0.6) mm, soft. |
subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
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Leaves | distal leaf sheaths persistent or disintegrating, pale brown, green or red-brown, streaked or mottled red-brown or purple, translucent, membranous, apex narrowly acute. |
distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
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Spikelets | basal spikelets absent; often proliferous, ovoid to ellipsoid or lanceoloid, terete, 2–10.7 × 1–2 mm, apex acute; proximal scale empty, persistent, amplexicaulous, dissimilar to floral scales, often longer, often resembling an involucral bract, ovate to lanceolate, midrib markedly thickened and broad and often prolonged beyond lamina; subproximal scale with flower; floral scales spiraled, colorless, whitish, or pale brown, streaked or mottled red-brown or purple, midribs green and red-brown or green and purple, ovate to elliptic, 0.8–1.5 × 0.4–0.8 mm, membranous, midrib obscure to prominent. |
ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
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Flowers | perianth bristles present or sometimes apparently absent; stamens 3; anthers 0.15–0.35; styles 3-fid. |
perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
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Achenes | whitish to olive or pale brown, sometimes spotted olive or red-brown, obovoid, trigonous (or subterete), angles prominent, 0.55–0.8 × 0.3–0.5 mm, apex constricted proximal to tubercle, smooth. |
falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
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Tubercles | green or pale brown to red-brown, trigonous. |
whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
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Eleocharis microcarpa |
Eleocharis bolanderi |
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Phenology | Fruiting late spring–summer. | |||||
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | |||||
Elevation | 1000–3400 m (3300–11200 ft) | |||||
Distribution |
AL; FL; GA; IN; LA; MA; MD; MI; MS; NC; NJ; PA; SC; TN; TX; VA; West Indies
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CA; CO; ID; NV; OR; UT
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Discussion | Varieties 2 (2 in the flora). Specimens of Eleocharis brittonii and E. microcarpa without achenes are often difficult to distinguish. More detailed study of the four apparent entities across the two species is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 95. | FNA vol. 23. | ||||
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. montevidensis var. bolanderi | |||||
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York 3: 312. (1836) | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | ||||
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