Eleocharis melanocarpa |
Eleocharis tuberculosa |
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black-fruit spike-rush, black-fruit spikesedge |
cone-cup spikerush, long-tubercled spikerush, long-tubercled spikesedge |
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Habit | Plants perennial, forming dense clumps, not stoloniferous. | |
Rhizomes | present or not, caudexlike, ascending, 1 mm thick; internodes and scales hidden by crowded culms, not evident. |
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Culms | to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular. |
elliptic or sometimes circular, with several rounded ridges and sulcate when dry, 15–75 cm × 0.3(–1.5) mm, firm, minutely granular at 10X. |
Leaves | distal leaf sheaths brown to reddish. |
distal leaf sheaths persistent; stramineous to green, often minutely red-spotted, papery to membranous, apex subacute to narrowly acute, often with tooth-like callus. |
Spikelets | 3–12 × 3–4 mm; proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire; subproximal scale with a flower; floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded. |
basal spikelets absent; never proliferous, ovoid, terete, 5–15 × 2.5–4 mm, apex acute; proximal scale empty, deciduous, clasping 1/2 of culm, similar to floral scales; subproximal scale often empty; floral scales spiraled, 10–30, 5–6 per mm of rachilla, pale orange-brown, midribs green, broadly ovate, 2.5–4 × 1.5–2.5 mm, central area and often flanks papery to cartilaginous (or membranous), midrib evident, apex broadly rounded to obtuse. |
Flowers | perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose; anthers brown, 1.7–2.2 mm. |
perianth bristles 5–6, bright brown, stout, slightly shorter than to equaling tubercle; spinules rarely absent, dense, variably from divaricate to antrorse, very short, and blunt to acute, to often much longer and retrorse, sharply acute; stamens 2–3; anthers 0.5–1 mm; styles 3-fid. |
Achenes | subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate. |
medium brown, obovoid to obpyriform, compressed (often obscurely) trigonous, angles evident, 0.9–1.7 × 0.8–2 mm; apex often constricted proximal to tubercle, very coarsely cancellate at 10X, each face with 8–10 longitudinal rows of large depressions. |
Tubercles | sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm. |
bone-white to pale orange-brown, often red-spotted, subpyramidal, cross section plano-convex, 0.9–1.7(–2.4) × 0.7–1.2–2(–2.2) mm, at least as high and wide as achene, spongy, apex rounded, often with an abaxial acute projection. |
Eleocharis melanocarpa |
Eleocharis tuberculosa |
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Phenology | Fruiting summer. | Fruiting spring (in south)–fall. |
Habitat | Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches | Wet soil, freshwater, ponds, lakeshores, streams, meadows, pine woods, grasslands, disturbed places, bogs |
Elevation | 10–300 m (0–1000 ft) | 0–500 m (0–1600 ft) |
Distribution |
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
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AL; AR; FL; GA; LA; MA; MS; NC; NJ; SC; TN; TX; NS
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Discussion | Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis tuberculosa evidently is closely related to E. tortilis, from which it differs in its larger tubercles, culms that are always elliptic in cross section, and perianth bristle spinules that are often divaricate to antrorse and often blunt. The specimens of E. tuberculosa from throughout its range which are like E. tortilis in having retrorsely spinulose perianth bristles have been named E. tuberculosa forma retrorsa Svenson (type from Massachusetts). Plants from Nova Scotia with nearly smooth bristles have been named E. tuberculosa var. pubnicoensis Fernald and E. tuberculosa forma pubnicoensis (Fernald) Svenson. We have not seen voucher specimens for literature reports of Eleocharis tuberculosa from Connecticut, Delaware, Maine, Maryland, New Hampshire, New York, Pennsylvania, Rhode Island, or Virginia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 89. | FNA vol. 23, p. 92. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus tuberculosus, E. tuberculosa var. pubnicoensis | |
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) | (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 152. (1817) |
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