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black-fruit spike-rush, black-fruit spikesedge

Parish spikerush, Parish's spike rush

Habit Plants perennial, mat-forming; rhizomes evident, long, 0.5–1(–2) mm thick, firm, cortex persistent, longer internodes 5–30 mm, scales often fugaceous, 5–10 mm, membranous, not fibrous.
Culms

to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular.

terete or cross section elliptic (or rectangular), usually with to 8 blunt ridges when dry, 10–50 cm × 0.2–0.7(–1) mm, soft to firm, spongy.

Leaves

distal leaf sheaths brown to reddish.

distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery to thickly membranous, apex usually redbrown, subtruncate to obtuse or subacute, callose, tooth often present on most or all culms, to 1 mm.

Spikelets

3–12 × 3–4 mm;

proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire;

subproximal scale with a flower;

floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded.

narrowly lanceoloid (to narrowly cylindric), 3–20 × 1.5–2.5 mm, apex acute;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed or spreading in fruit, 15–40, 3–4 per mm of rachilla, orange brown, midrib regions often greenish, ovate, 2–3 × 1 mm, apex entire, rounded to obtuse in proximal part of spikelet, acute in distal part, carinate, at least in distal part of spikelet.

Flowers

perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose;

anthers brown, 1.7–2.2 mm.

perianth bristles 3–7, rarely apparently absent, stramineous to whitish, slender to stout, often unequal, rudimentary to slightly exceeding tubercle;

stamens 3;

anthers yellow to stramineous, 1.1–2 mm;

styles 3-fid, 0.3–0.4 × 0.2–0.3 mm.

Achenes

subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate.

falling with scales, yellow-brown or dark brown, ellipsoid or obovoid to obpyriform, compressed-trigonous, angles evident to obscure, rarely prominent, 0.8–1.4 × 0.5–0.7 mm, neck absent to long, smooth or sometimes minutely cancellate at 20–30X.

Tubercles

sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm.

whitish, pyramidal, often higher than wide, 0.3–0.4 × 0.25–0.35 mm.

2n

= 10.

Eleocharis melanocarpa

Eleocharis parishii

Phenology Fruiting summer. Fruiting spring–fall.
Habitat Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches Fresh to brackish, wet soil or emergent, often drying lakeshores, ponds, streams, springs
Elevation 10–300 m (0–1000 ft) 500–2300 m (1600–7500 ft)
Distribution
from FNA
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; KS; NM; NV; OR; TX; UT; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis parishii is very similar to E. montevidensis, with which it sometimes grows. It differs from E. montevidensis mainly in its narrowly lanceoloid to cylindric spikelets and its floral scale apices not recurved or horizontally wrinkled. Some apparent intermediates occur in regions of sympatry. The floral scales of E. montevidensis are more densely placed on the rachilla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 89. FNA vol. 23, p. 80.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. disciformis, E. montevidensis var. disciformis, E. montevidensis var. parishii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) Britton: J. New York Microscop. Soc. 5: 110. (1889)
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