Eleocharis melanocarpa |
Eleocharis pachycarpa |
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black-fruit spike-rush, black-fruit spikesedge |
black sand spikerush, broad fruit spikerush |
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Habit | Plants perennial, forming dense clumps, often stoloniferous. | |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
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Culms | to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular. |
erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
Leaves | distal leaf sheaths brown to reddish. |
distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
Spikelets | 3–12 × 3–4 mm; proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire; subproximal scale with a flower; floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded. |
basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
Flowers | perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose; anthers brown, 1.7–2.2 mm. |
perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate. |
stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
Tubercles | sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm. |
stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
Eleocharis melanocarpa |
Eleocharis pachycarpa |
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Phenology | Fruiting summer. | Fruiting late spring–summer. |
Habitat | Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches | Fresh shores, streambeds, groundwater seeps |
Elevation | 10–300 m (0–1000 ft) | 100–1400 m (300–4600 ft) |
Distribution |
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
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CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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Discussion | Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 89. | FNA vol. 23, p. 91. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Name authority | Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) |
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