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black-fruit spike-rush, black-fruit spikesedge

neat spike-rush, quill spike-rush, quill spikesedge, slender spikerush, éléocharide brillante

Habit Plants perennial, mat-forming; rhizomes evident, 0.3–0.5 mm thick, hard, cortex persistent, longer internodes 2 mm, scales persistent or fugaceous, 2–3 mm, membranous to papery, slightly fibrous.
Culms

to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular.

terete, 2–15 cm × 0.15–0.3 mm, firm to soft.

Leaves

distal leaf sheaths brown to reddish.

distal leaf sheaths persistent, not splitting, proximally stramineous to reddish, distally green or stramineous to reddish, membranous, apex often red, obtuse to acute, tooth absent.

Spikelets

3–12 × 3–4 mm;

proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire;

subproximal scale with a flower;

floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded.

ovoid, 1–4 × 1–2 mm, obtuse to acute;

proximal scale amplexicaulous, apex entire;

subproximal scale with flower;

floral scales spreading in fruit, 5–30, 8 per mm of rachilla, medium to very dark brown, midrib region often pale or greenish, broadly ovate, 1–1.3 × 1 mm, apex rounded, entire, not carinate.

Flowers

perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose;

anthers brown, 1.7–2.2 mm.

perianth bristles absent;

stamens 3;

anthers yellow, 0.3 mm;

styles 3-fid.

Achenes

subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate.

persistent after scales fall, to dark yellow-orange or brown, broadly obpyriform, trigonous, angles evident, 0.6–0.65 × 0.5–0.55 mm, rugulose at 20–30X, 20 blunt horizontal ridges in each vertical series.

Tubercles

sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm.

brown, greatly depressed, rudimentary, 0.05–15 × 0.15–0.3 mm.

Eleocharis melanocarpa

Eleocharis nitida

Phenology Fruiting summer. Fruiting late spring (Jun)–summer.
Habitat Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches Fresh bog pools, streams, disturbed places
Elevation 10–300 m (0–1000 ft) 30–400 m (100–1300 ft)
Distribution
from FNA
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MI; MN; NH; WI; NL; NS; ON; QC; SK
[BONAP county map]
Discussion

Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis nitida is much like E. elliptica but all structures are smaller; intermediates are unknown.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 89. FNA vol. 23, p. 86.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) Fernald: Rhodora 8: 129. (1906)
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