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black-fruit spike-rush, black-fruit spikesedge

slim spikerush

Habit Plants perennial; rhizomes 1–1.5 mm thick, soft, longer internodes 2–3 cm, scales 5–14 mm, tubers absent.
Culms

to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular.

obscurely trigonous to terete;

spikelet-bearing culms 16–80 cm × 0.5–1.5 mm; when submersed plants often forming numerous, filiform flaccid culms without spikelets, sometimes with whorls of slender branches, 0.1–0.3 mm wide, soft;

sometimes septate-nodulose when aquatic, internally spongy, transverse septa incomplete.

Leaves

distal leaf sheaths brown to reddish.

distal leaf sheaths persistent or decaying, membranous, apex acute, often prolonged into translucent portion to 1 mm.

Spikelets

3–12 × 3–4 mm;

proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire;

subproximal scale with a flower;

floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded.

not proliferous, (6–)9–24 × 1.4–2.2 mm;

rachilla joints bearing prominent winglike remnants of floral scales;

proximal scale with a flower, amplexicaulous, 2.5–4.1 mm;

floral scales 7–26, 1–2 per mm of rachilla, green to stramineous or pale brown, often minutely dotted reddish, usually with conspicuous dark brown to blackish submarginal band, narrowly ovate, 3.5–4.5 × 2 mm, thickly papery, membranous toward margins.

Flowers

perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose;

anthers brown, 1.7–2.2 mm.

perianth bristles 6–7, whitish to stramineous or pale reddish brown, proximally slightly flattened, unequal, exceeding or rarely shorter than achene, 0.7–1.9 mm, retrorsely spinulose;

anthers yellow to reddish, 1.7–1.9 mm;

styles 3-fid.

Achenes

subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate.

whitish, stramineous, or pale green, obovoid to obpyriform, compressed trigonous with adaxial face broadest, or biconvex, 0.65–1.4 × 0.5–0.8 mm, clearly sculptured at 10–15X, each face with 10–13 rows of rectangular, transversely elongated cells, apex constricted to short neck 0.2–0.25(–0.3) mm wide, wider at tubercle base.

Tubercles

sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm.

dark brown, pyramidal, 0.2–0.5 × 0.2–0.4 mm.

Eleocharis melanocarpa

Eleocharis elongata

Phenology Fruiting summer. Fruiting late spring–late fall.
Habitat Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches Sometimes drying ponds, lakeshores, marshes, creeks, canals, ditches
Elevation 10–300 m (0–1000 ft) 10–50 m (0–200 ft)
Distribution
from FNA
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; NC; TX; Mexico; Central America; South America
[BONAP county map]
Discussion

Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis elongata sometimes grows with E. robbinsii; no intermediates are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 89. FNA vol. 23, p. 117.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) Chapman: Fl. South. U.S., 515. (1860)
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