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black-fruit spike-rush, black-fruit spikesedge

two-tone spikerush

Habit Plants probably perennial, tufted, not stoloniferous; rhizomes present, ± horizontal, 0.2–0.4 mm thick, firm, longer internodes 3–7 mm, scales translucent or not.
Culms

to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular.

often ascending, often spotted or streaked red-brown, cross section quadrangular or broadly elliptic, often sulcate, 1.5–8(–25) cm × 0.1–0.3 mm, soft.

Leaves

distal leaf sheaths brown to reddish.

distal leaf sheaths persistent or fugaceous, closely sheathing, not conspicuously wrinkled, colorless or pale brown, sometimes spotted red-brown, delicately membranous, apex narrowly acute.

Spikelets

3–12 × 3–4 mm;

proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire;

subproximal scale with a flower;

floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded.

basal spikelets absent; never proliferous, ovoid or ellipsoid, laterally compressed or terete, 1.5–3.7 × 0.8–3 mm, apex acute;

proximal scale empty, persistent, amplexicaulous, similar to floral scales;

subproximal scale with a flower;

floral scales distichous (spirodistichous), 8–22, 6–10 per mm of rachilla, colorless or pale brown, mottled red-brown to purple, ovate, (1–)2 × 1 mm, membranous, midribs green and red-brown, prominent, apex rounded to obtuse.

Flowers

perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose;

anthers brown, 1.7–2.2 mm.

perianth bristles present or absent, (1–)4, colorless to white, vestigial to shorter than achene, 0–0.55 mm;

spinules not evident at 45X;

stamens (1–)3;

anthers 0.35–0.45 mm;

styles 2-fid (or 3-fid).

Achenes

subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate.

whitish to pale brown, obovoid, biconvex, rarely trigonous, angles prominent, 0.5–0.8 × 0.4–0.6 mm, apex slightly constricted proximal to tubercle, smooth.

Tubercles

sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm.

sessile, green or red-brown, pyramidal, birettaform or umbonate, 0.15–0.25 × 0.3–0.4 mm.

Eleocharis melanocarpa

Eleocharis bicolor

Phenology Fruiting summer. Fruiting summer–late fall.
Habitat Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches Moist banks, freshwater, damp pine barrens
Elevation 10–300 m (0–1000 ft) 0–50 m (0–200 ft)
Distribution
from FNA
AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; GA; MS; West Indies; Central America (Nicaragua)
Discussion

Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The culms of Eleocharis bicolor sometimes have complete transverse septa which are prominent when plants are aquatic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 89. FNA vol. 23, p. 96.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) Chapman: Fl. South. U.S., 517. (1860)
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