Eleocharis interstincta |
Eleocharis quadrangulata |
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knotted spikerush |
four-angle spikerush, square-stem spike-rush, square-stem spikesedge |
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Habit | Plants perennial; rhizomes 2–4 mm thick, firm, longer internodes 3–7.5 cm, scales 5 mm, tubers absent. | Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent. |
Culms | terete, 45–100 cm × (3.2–)5–9.4 mm, soft to firm, sometimes septate-nodulose proximally, not distally, internally hollow with complete transverse septa, closer together near the spikelet, evident externally; plants never forming filiform, flaccid culms. |
acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete; plants never forming filiform, flaccid culms. |
Leaves | distal leaf sheaths persistent, membranous to thinly papery, apex acute to acuminate. |
distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm. |
Spikelets | not proliferous, (20–)40–62 × 4–7 mm; rachilla joints bearing prominent wing-like remnants of floral scales; proximal scale empty, amplexicaulous, (3–)3.5–5 mm; floral scales 115–220, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, obovate to broadly oblong, (4–)4.5–5 × 2.8–4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to subacute. |
not proliferous, (15–)20–76 × 3–5(–6) mm; rachilla joints bearing obscure winglike remnants of floral scales; proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm; floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse. |
Flowers | perianth bristles 6–8, stramineous, stout, flattened, subequal, exceeding achene, to 2.9 mm, coarsely spinulose; anthers stramineous to reddish, 2.5–5 mm; styles 2-fid or 3-fid. |
perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth; anthers stramineous to red-brown, 2.3–2.9 mm; styles 3-fid, sometimes 2-fid. |
Achenes | stramineous to golden-yellow or reddish brown or gray, obovoid to obpyriform, biconvex or nearly plano-convex, often with abaxial longitudinal ridge, 1.4–1.8(–2) × 1.1–1.4 mm, markedly sculptured at 10–15X, each face with 23–37 rows of transversely elongated cells, the longitudinal walls separating the cells often prominent, apex with short neck 0.7–0.8 mm wide. |
yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide. |
Tubercles | dark brown, lamelliform, slightly higher than wide, 0.7–1.1 × 0.5–0.7 mm. |
dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy. |
Eleocharis interstincta |
Eleocharis quadrangulata |
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Phenology | Fruiting late spring–winter. | Fruiting early summer–winter. |
Habitat | Fresh ponds, lakeshores, marshes, springs, ditches, canals | Shallow water of fresh lake and pond shores, marshes |
Elevation | 10–500 m (0–1600 ft) | 10–600 m (0–2000 ft) |
Distribution |
FL; OK; TX; Mexico; Central America; e South America
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AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
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Discussion | Contrary to statements in the literature, sectioning reveals that the culm septa are closer together near the spikelet than in the rest of the culm in both Eleocharis interstincta and E. equisetoides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee. The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 119. | FNA vol. 23, p. 120. |
Parent taxa | Cyperaceae > Eleocharis > subg. Limnochloa | Cyperaceae > Eleocharis > subg. Limnochloa |
Sibling taxa | ||
Synonyms | Scirpus interstinctus | Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus |
Name authority | (Vahl) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 149. (1817) | (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817) |
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