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knotted spikerush

Parish spikerush, Parish's spike rush

Habit Plants perennial; rhizomes 2–4 mm thick, firm, longer internodes 3–7.5 cm, scales 5 mm, tubers absent. Plants perennial, mat-forming; rhizomes evident, long, 0.5–1(–2) mm thick, firm, cortex persistent, longer internodes 5–30 mm, scales often fugaceous, 5–10 mm, membranous, not fibrous.
Culms

terete, 45–100 cm × (3.2–)5–9.4 mm, soft to firm, sometimes septate-nodulose proximally, not distally, internally hollow with complete transverse septa, closer together near the spikelet, evident externally;

plants never forming filiform, flaccid culms.

terete or cross section elliptic (or rectangular), usually with to 8 blunt ridges when dry, 10–50 cm × 0.2–0.7(–1) mm, soft to firm, spongy.

Leaves

distal leaf sheaths persistent, membranous to thinly papery, apex acute to acuminate.

distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery to thickly membranous, apex usually redbrown, subtruncate to obtuse or subacute, callose, tooth often present on most or all culms, to 1 mm.

Spikelets

not proliferous, (20–)40–62 × 4–7 mm;

rachilla joints bearing prominent wing-like remnants of floral scales;

proximal scale empty, amplexicaulous, (3–)3.5–5 mm;

floral scales 115–220, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, obovate to broadly oblong, (4–)4.5–5 × 2.8–4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to subacute.

narrowly lanceoloid (to narrowly cylindric), 3–20 × 1.5–2.5 mm, apex acute;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed or spreading in fruit, 15–40, 3–4 per mm of rachilla, orange brown, midrib regions often greenish, ovate, 2–3 × 1 mm, apex entire, rounded to obtuse in proximal part of spikelet, acute in distal part, carinate, at least in distal part of spikelet.

Flowers

perianth bristles 6–8, stramineous, stout, flattened, subequal, exceeding achene, to 2.9 mm, coarsely spinulose;

anthers stramineous to reddish, 2.5–5 mm;

styles 2-fid or 3-fid.

perianth bristles 3–7, rarely apparently absent, stramineous to whitish, slender to stout, often unequal, rudimentary to slightly exceeding tubercle;

stamens 3;

anthers yellow to stramineous, 1.1–2 mm;

styles 3-fid, 0.3–0.4 × 0.2–0.3 mm.

Achenes

stramineous to golden-yellow or reddish brown or gray, obovoid to obpyriform, biconvex or nearly plano-convex, often with abaxial longitudinal ridge, 1.4–1.8(–2) × 1.1–1.4 mm, markedly sculptured at 10–15X, each face with 23–37 rows of transversely elongated cells, the longitudinal walls separating the cells often prominent, apex with short neck 0.7–0.8 mm wide.

falling with scales, yellow-brown or dark brown, ellipsoid or obovoid to obpyriform, compressed-trigonous, angles evident to obscure, rarely prominent, 0.8–1.4 × 0.5–0.7 mm, neck absent to long, smooth or sometimes minutely cancellate at 20–30X.

Tubercles

dark brown, lamelliform, slightly higher than wide, 0.7–1.1 × 0.5–0.7 mm.

whitish, pyramidal, often higher than wide, 0.3–0.4 × 0.25–0.35 mm.

2n

= 10.

Eleocharis interstincta

Eleocharis parishii

Phenology Fruiting late spring–winter. Fruiting spring–fall.
Habitat Fresh ponds, lakeshores, marshes, springs, ditches, canals Fresh to brackish, wet soil or emergent, often drying lakeshores, ponds, streams, springs
Elevation 10–500 m (0–1600 ft) 500–2300 m (1600–7500 ft)
Distribution
from FNA
FL; OK; TX; Mexico; Central America; e South America
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from FNA
AZ; CA; KS; NM; NV; OR; TX; UT; Mexico
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[BONAP county map]
Discussion

Contrary to statements in the literature, sectioning reveals that the culm septa are closer together near the spikelet than in the rest of the culm in both Eleocharis interstincta and E. equisetoides.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis parishii is very similar to E. montevidensis, with which it sometimes grows. It differs from E. montevidensis mainly in its narrowly lanceoloid to cylindric spikelets and its floral scale apices not recurved or horizontally wrinkled. Some apparent intermediates occur in regions of sympatry. The floral scales of E. montevidensis are more densely placed on the rachilla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 119. FNA vol. 23, p. 80.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus interstinctus E. disciformis, E. montevidensis var. disciformis, E. montevidensis var. parishii
Name authority (Vahl) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 149. (1817) Britton: J. New York Microscop. Soc. 5: 110. (1889)
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