Eleocharis geniculata |
Eleocharis coloradoensis |
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bent spike-rush, Canada spikesedge, capitate spike-rush |
dwarf spike-rush |
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Habit | Plants tufted, without creeping rhizomes. | |
Culms | to 45 cm × 0.2–1 mm. |
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Leaves | distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute. |
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Spikelets | orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute; proximal scale without flower, not amplexicaulous; floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute. |
3–6 × 1–1.5 mm; proximal scale 1/2 or less of spikelet length; floral scales 6–25 per spikelet, mostly bright orange-brown, sometimes stramineous, 1.7–2.5 mm, apex subacute. |
Flowers | perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense; styles 2-fid. |
perianth bristles mostly absent or rudimentary, occasionally to 5, 1/2 of achene length, very unequal; anthers 0.6–0.9 mm. |
Achenes | brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X. |
medium to dark brown, obovoid to obpyriform, thickly trigonous, rarely some biconvex, angles distinct to prominent, faces convex or some plane, 0.75–1.1 × 0.55–0.7 mm, apex narrowly truncate or tapered into tubercle, rugulose at 10X to finely rough at 30X. |
Tubercles | stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute. |
0.05–0.2 × 0.15 mm. |
Tubers | terminating rhizomes, 2.5–4 × 0.7–1.5 mm, body (apart from apical bud) broadly oblong to orbicular, not markedly curved. |
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2n | = 10. |
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Eleocharis geniculata |
Eleocharis coloradoensis |
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Phenology | Fruiting spring–winter (Mar–Dec). | Fruiting summer–fall (north), spring–fall (far south). |
Habitat | Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes | Fresh or brackish drying lake and pond margins, stream beds, flood plains, vernal pools, irrigation ditches, tidal wetlands |
Elevation | 0–1500 m [0–4900 ft] | 0–2100 m [0–6900 ft] |
Distribution |
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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AL; AR; CA; CO; IA; ID; KS; LA; MN; MO; MS; ND; NM; NV; OK; OR; SD; TX; UT; MB; SK; Mexico |
Discussion | The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most authors, except C. L. Gilly (1941), H. L. Mason (1957), and R. R. Yeo (1980), have included Eleocharis coloradoensis in E. parvula or E. parvula var. anachaeta. In typical E. coloradoensis, which occurs from Saskatchewan south to Kansas and in California to 2100 m elevation, the achenes are usually distinctly rugulose or rough, often pitted-cellular, their apices usually truncate, and the tubercles are usually brown, often rudimentary, clearly distinct from the achene, and apparently partly sunken into the achene summit. Some plants from the southern Great Plains to the Mexican border, including the type of E. parvula var. anachaeta from Louisiana, may deserve taxonomic recognition. They differ from typical E. coloradoensis in having nearly smooth achenes with the apex tapered to a tubercle that is difficult to distinguish from the achene. C. L. Gilly (1941) separated these plants as E. membranacea (Buckley) Gilly; application of that name is doubtful because achenes are lacking from the type. R. R. Yeo (1980) studied the life-history of E. coloradoensis in the Sacramento Valley, California, and showed that it can be used to control several aquatic weeds in irrigation canals. The n = 4 count reported from Kansas under E. parvula var. anachaeta (Anonymous 1964) and on voucher specimens at GH and UC, is probably erroneous; the label on a duplicate voucher specimen at NDA includes the information “n = 3 II’s + a chain of IV” (i.e., n = 5). The record from Washington is somewhat doubtful because the specimen lacks achenes. Literature reports of E. parvula from Illinois and Tennessee may refer to E. coloradoensis; I have not seen specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 101. | FNA vol. 23, p. 106. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Scirpus geniculatus, E. capitata, E. caribaea, E. dispar | Scirpus coloradoensis, E. parvula var. anachaeta |
Name authority | (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) | (Britton) Gilly: Amer. Midl. Naturalist 26: 66. (1941) |
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