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bent spike-rush, Canada spikesedge, capitate spike-rush

cone-cup spikerush, long-tubercled spikerush, long-tubercled spikesedge

Habit Plants tufted, without creeping rhizomes. Plants perennial, forming dense clumps, not stoloniferous.
Rhizomes

present or not, caudexlike, ascending, 1 mm thick;

internodes and scales hidden by crowded culms, not evident.

Culms

to 45 cm × 0.2–1 mm.

elliptic or sometimes circular, with several rounded ridges and sulcate when dry, 15–75 cm × 0.3(–1.5) mm, firm, minutely granular at 10X.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent; stramineous to green, often minutely red-spotted, papery to membranous, apex subacute to narrowly acute, often with tooth-like callus.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

basal spikelets absent; never proliferous, ovoid, terete, 5–15 × 2.5–4 mm, apex acute;

proximal scale empty, deciduous, clasping 1/2 of culm, similar to floral scales;

subproximal scale often empty;

floral scales spiraled, 10–30, 5–6 per mm of rachilla, pale orange-brown, midribs green, broadly ovate, 2.5–4 × 1.5–2.5 mm, central area and often flanks papery to cartilaginous (or membranous), midrib evident, apex broadly rounded to obtuse.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 5–6, bright brown, stout, slightly shorter than to equaling tubercle;

spinules rarely absent, dense, variably from divaricate to antrorse, very short, and blunt to acute, to often much longer and retrorse, sharply acute;

stamens 2–3;

anthers 0.5–1 mm;

styles 3-fid.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

medium brown, obovoid to obpyriform, compressed (often obscurely) trigonous, angles evident, 0.9–1.7 × 0.8–2 mm;

apex often constricted proximal to tubercle, very coarsely cancellate at 10X, each face with 8–10 longitudinal rows of large depressions.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

bone-white to pale orange-brown, often red-spotted, subpyramidal, cross section plano-convex, 0.9–1.7(–2.4) × 0.7–1.2–2(–2.2) mm, at least as high and wide as achene, spongy, apex rounded, often with an abaxial acute projection.

2n

= 10.

Eleocharis geniculata

Eleocharis tuberculosa

Phenology Fruiting spring–winter (Mar–Dec). Fruiting spring (in south)–fall.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Wet soil, freshwater, ponds, lakeshores, streams, meadows, pine woods, grasslands, disturbed places, bogs
Elevation 0–1500 m (0–4900 ft) 0–500 m (0–1600 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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from FNA
AL; AR; FL; GA; LA; MA; MS; NC; NJ; SC; TN; TX; NS
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Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis tuberculosa evidently is closely related to E. tortilis, from which it differs in its larger tubercles, culms that are always elliptic in cross section, and perianth bristle spinules that are often divaricate to antrorse and often blunt. The specimens of E. tuberculosa from throughout its range which are like E. tortilis in having retrorsely spinulose perianth bristles have been named E. tuberculosa forma retrorsa Svenson (type from Massachusetts). Plants from Nova Scotia with nearly smooth bristles have been named E. tuberculosa var. pubnicoensis Fernald and E. tuberculosa forma pubnicoensis (Fernald) Svenson.

We have not seen voucher specimens for literature reports of Eleocharis tuberculosa from Connecticut, Delaware, Maine, Maryland, New Hampshire, New York, Pennsylvania, Rhode Island, or Virginia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23, p. 92.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar Scirpus tuberculosus, E. tuberculosa var. pubnicoensis
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 152. (1817)
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