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bent spike-rush, Canada spikesedge, capitate spike-rush

Habit Plants tufted, without creeping rhizomes. Plants perennial, densely tufted; rhizomes mostly hidden by aerial shoots and roots, not long, 2–3 mm thick, hard, cortex persistent?, internodes very short, scales persistent, 4–7 mm, membranous, slightly fibrous.
Culms

to 45 cm × 0.2–1 mm.

terete, when dry with to 12 blunt ribs, 20–55 cm × 0.5–1.1 mm, soft to firm, internally mostly hollow with complete transverse septa 2–4 mm apart, evident only on sectioning culm.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, not splitting, proximally dark red, distally stramineous or reddish, thinly papery, apex often red to brown, obtuse to subtruncate, slightly callose, tooth present, 0.5–1(–3.7) mm.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

lanceoloid, 5–13 × 2–2.5 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale empty;

floral scales appressed in fruit, 10–100, 10–12 per mm of rachilla, medium brown to stramineous, midrib regions often greenish, ovate, 1–1.5 × 1 mm, entire, apex rounded to subacute, carinate in distal part of spikelet.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 5–6, pale brown, stout, equaling achene;

stamens 3;

anthers brown, 0.4–0.9 mm;

styles 3-fid or a few 2-fid in the same spikelet.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

falling with scales, green or medium or dark brown, obpyriform, compressed-trigonous or some biconvex in same spikelet, angles prominent, 0.6–0.8 × 0.5–0.6 mm, neck short or absent, smooth or very finely reticulate at 20–30X.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

brown, depressed-pyramidal, 0.1–0.2 × 0.2–0.3 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis ravenelii

Phenology Fruiting spring–winter (Mar–Dec). Fruiting spring–fall.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Fresh, wet to damp, seasonally wet depressions, flatwoods, ditches
Elevation 0–1500 m (0–4900 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (San Luis Potosí)
[BONAP county map]
Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis ravenelii is apparently very uncommon in North America. It is often mistaken for very slender-stemmed E. montana, which differs in its spikelets with floral scales 1.5 mm or more, 100–500 per spikelet and 15–40 per mm of rachilla, its mostly larger and biconvex achenes, and its culm septa usually evident without sectioning the culm. Although the holotype of E. ravenelii (NY), from Corpus Christi, Texas, lacks culm bases and leaf sheaths, its culms, spikelets, floral scales, and achenes are typical of the later E. austrotexana M. C. Johnston.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar E. austrotexana
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) Britton: in J. K. Small, Fl. S.E. U.S., 184, 1327. (1903)
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