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bent spike-rush, Canada spikesedge, capitate spike-rush

spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial; rhizomes 3–4 mm thick, soft to hard, longer internodes 3–8 cm, scales more than 1 cm, tubers absent.
Culms

to 45 cm × 0.2–1 mm.

obscurely 3–5-angled to terete, 40–110 cm × (2–)3.6–7.5 mm, soft, not septate-nodulose, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, membranous, apex acute to long-acuminate, sometimes prolonged into a slender bladelike portion to 6 cm.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

not proliferous, 12–43 × 3.5–6 mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale without a flower, amplexicaulous, 3.6–7.5 mm;

floral scales 30–125, 1–3 per mm of rachilla, greenish to pale brown, often with pale to dark brown submarginal band or a subapical darker spot, ovate to oblong, 4.3–5.8 × 2.5–3.3 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to acute.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 6–7, stramineous, margins and spinules reddish to pale brown, stout, flattened, subequal, exceeding achene, 2.5–3.2 mm, coarsely retrorsely spinulose;

anthers reddish brown, 1.5–1.9 mm;

styles 3-fid or sometimes 2-fid.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

pale brown, biconvex, obpyriform, 1.7–2.3 × 1.2–1.5 mm, clearly sculptured at 10–15X, each face with 10–15 rows of transversely elongated cells, often isodiametric at achene base, apex usually constricted to neck 0.5–0.6 mm wide.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

stramineous, lamelliform to high-pyramidal, 0.8–1.1 × 0.7–0.9 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis obtusetrigona

Phenology Fruiting spring–winter (Mar–Dec). Fruiting spring–fall.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Fresh, permanent water in ditches
Elevation 0–1500 m (0–4900 ft) 0–20 m (0–100 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; e Mexico; South America; Central America (Nicaragua)
[BONAP county map]
Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In North America Eleocharis obtusetrigona is known from only five collections, two of them originally identified as E. fistulosa (Poiret) Link [= E. acutangula (Roxburgh) Schultes], which is common in Mexico and South America. The original description of E. obtusetrigona is poor and does not clearly distinguish E. obtusangula from E. acutangula. H. K. Svenson (1939) included E. obtusetrigona in E. acutangula, which can be distinguished by its acutely trigonous culms as well as its smaller floral scales and achenes. The tubercles are often spongy as in E. quadrangulata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23, p. 118.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar Limnochloa obtusetrigona, E. fistulosa var. obtusetrigona, E. mutata var. obtusetrigona
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) (Lindley & Nees) Steudel: Syn. Pl. Glumac. 2: 80. (1855)
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