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bent spike-rush, Canada spikesedge, capitate spike-rush

mountain spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial, densely tufted or mat-forming; rhizomes mostly hidden by culms and roots, fairly long, 3 mm thick, hard, cortex persistent, longer internodes to 3 mm, scales persistent, ca. 8 mm, membranous, slightly fibrous.
Culms

to 45 cm × 0.2–1 mm.

terete, when dry with few to many blunt ridges, 10–70 cm × 0.5–2 mm, soft to firm, internally hollow with complete transverse septa 2–5 mm apart, usually evident externally except in narrowest culms.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, not splitting, proximally dark red, distally green or brown, slightly callose, papery, apex obtuse, tooth present, 0.3–1 mm.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

ovoid, 6–21 × 3–4 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 100–500+, (15–)30–40 per mm of rachilla, medium brown to colorless, midrib regions greenish to colorless, ovate, 1.5–2.5 × 1–1.5 mm, entire, apex rounded to subacute, carinate in distal part of spikelet.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 6–8, pale brown, medium stout, from less than 1/2 achene length to sometimes slightly exceeding tubercle, retrorsely spinulose;

stamens 1;

anthers dark yellow to brown, 0.6–1 mm;

styles 3-fid or some 2-fid.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

falling with scales, dark green or medium brown, obovoid to obpyriform, biconvex or sometimes some compressedtrigonous in same spikelet, lateral angles prominent, abaxial angle absent or evident, not prominent, 0.8–1.1 × 0.7–0.8 mm, neck short or absent, finely cancellate at 10–20X, sometimes finely rugulose, with 15 horizontal ridges in vertical series.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

brown, pyramidal, mostly depressed, 0.2–0.35 × 0.2–0.4 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis montana

Phenology Fruiting spring–winter (Mar–Dec). Fruiting winter–fall.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Fresh temporary or artificial ponds, ditches, burned savannas, swamp margins
Elevation 0–1500 m (0–4900 ft) 20–90[–2800] m (100–300[–9200] ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
AL; AZ; FL; GA; LA; TX; Mexico; Central America; South America; West Indies
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Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The taxonomy of the septate-culmed species here treated as Eleocharis montana and E. ravenelii should be evaluated. According to H. K. Svenson (1957), the type of E. montana from near Bogotá, Colombia, is the mountain extreme of the species; it has swollen culms with no visible septation.

Specimens from Acadia and St. Landry parishes, Louisiana, are intermediate between Eleocharis montana and E. montevidensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23, p. 79.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar Scirpus montanus, E. montana var. nodulosa, E. nodulosa
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) (Kunth) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 153. (1817)
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