Eleocharis geniculata |
Eleocharis fallax |
|
---|---|---|
bent spike-rush, Canada spikesedge, capitate spike-rush |
creeping spikerush, creeping spikesedge |
|
Habit | Plants tufted, without creeping rhizomes. | Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 5–10(–20) mm, scales sometimes fugaceous, 5–7 mm, papery (to membranous), sometimes fibrous. |
Culms | to 45 cm × 0.2–1 mm. |
terete, often with to 12 blunt ridges when dry, 30–75 cm × 0.5–1.5 mm, firm, internally spongy. |
Leaves | distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute. |
distal leaf sheaths persistent, not splitting, proximally dark red, distally stramineous or green, papery to membranous, apex dark redbrown, obtuse, not callose, tooth rarely present on some culms, 0.2 mm. |
Spikelets | orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute; proximal scale without flower, not amplexicaulous; floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute. |
ovoid or subspheric, 5–12 × 2–3(–4) mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales deciduous, often spreading in fruit, 10–35, 3 per mm of rachilla, red-brown to blackish brown, midrib regions sometimes green, ovate, 2.5–3 × 1.7 mm, apex entire, obtuse to acute, often carinate in distal part of spikelet. |
Flowers | perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense; styles 2-fid. |
perianth bristles 1–5, brown, stout, very unequal, rudimentary to equaling achene; stamens unknown [absent from specimens]; styles 3-fid or some 2-fid. |
Achenes | brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X. |
not persistent, dark yellow or medium brown, obovoid to obpyriform, compressed trigonous, or some thickly biconvex, angles evident, 1.1–1.5 × 0.95–1.25 mm, apex rounded, neck very short (to absent), finely rugulose at 10–20X, 20 or more horizontal ridges in vertical series, finely cancellate at 20–30X. |
Tubercles | stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute. |
whitish to brown, pyramidal, not depressed, as high as wide, 0.3–0.5 × 0.4–0.6 mm. |
2n | = 10. |
= 42. |
Eleocharis geniculata |
Eleocharis fallax |
|
Phenology | Fruiting spring–winter (Mar–Dec). | Fruiting late summer. |
Habitat | Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes | Coastal, fresh to brackish pond and lakeshores, marsh |
Elevation | 0–1500 m (0–4900 ft) | 0–10 m (0–0 ft) |
Distribution |
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
|
MA; NJ; NS |
Discussion | The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. The taxonomic status of Eleocharis fallax is problematic. I have seen only the type, from 1919 from a pond on Cape Cod, Massachusetts; a specimen from 1955 from Inverness County, Nova Scotia; and a specimen from the Passaic River marshes in New Jersey. It is reportedly extirpated from the type locality (M. L. Fernald 1950). Another specimen from Cape Cod is typical of E. fallax except for some lenticular achenes. I have not seen a voucher for the material, reportedly collected by H. K. Svenson from a brackish marsh on Plum Island, Essex County, Massachusetts, from which S.-O. Strandhede (1967) counted the chromosome number. It seems possible that these collections are of hybrid origin from E. elliptica and either E. erythropoda or E. uniglumis var. halophila (M. L. Fernald 1950). The plants are similar to E. erythropoda except for their trigonous, rougher achenes and sometimes fibrous rhizome scales, which suggests introgression from E. elliptica. Because some specimens of E. ambigens, including the type, have mixtures of lenticular and trigonous achenes, E. fallax is often treated as conspecific with E. ambigens, from which it differs only in its 3-fid styles, trigonous, more rugulose achenes, and higher tubercles. It seems best to recognize E. fallax as a species pending further research because the plants bear many, apparently normal achenes and are not exactly intermediate between their putative parents, and to avoid using a name for the widespread E. ambigens that may apply only to rare hybrids. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 101. | FNA vol. 23, p. 78. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | Scirpus geniculatus, E. capitata, E. caribaea, E. dispar | |
Name authority | (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) | Weatherby: Rhodora 24: 23. (1922) |
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