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bent spike-rush, Canada spikesedge, capitate spike-rush

horsetail spike-rush, horsetail spikesedge, jointed spike-rush, jointed spike-rush (spikesedge), jointed spikesedge, knotted spike-rush

Habit Plants tufted, without creeping rhizomes. Plants perennial; rhizomes 2–4 mm thick, soft to firm, longer internodes 2–8 cm, scales persistent, 7–8 mm, membranous, tubers absent.
Culms

to 45 cm × 0.2–1 mm.

terete, 50–100 cm × 5–8.4 mm, soft to firm, obscurely to clearly septate-nodulose throughout, internally hollow with complete transverse septa, closer together near the spikelet;

plants never forming filiform, flaccid culms.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, membranous to thinly papery, apex acute to slightly acuminate.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

not proliferous, 20–45 × 4–5(–8) mm;

rachilla joints bearing prominent winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (2–)2.8–4.9 mm;

floral scales 85–160, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, obovate to oblong, (4.8–)5.2–7 × 3–4.4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to obtuse.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 3–8, stramineous to reddish, slender, vestigial to 2.5 mm, shorter than to equaling achene, smooth or obscurely retrorsely spinulose;

anthers stramineous to red-striated, 2.6–3.2 mm;

styles 2–3-fid.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

yellowish to reddish brown, obovoid, biconvex to obscurely compressed trigonous, 1.8–2.3 × 1.4–1.9 mm, obscurely sculptured or sometimes partially smooth at 10–15X, each face with 20–40 rows of linear, transversely elongated cells, lines separating cells obscure, apex with very short neck 0.6–0.8 mm wide.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

dark brown, lamelliform, high-deltoid, 0.6–1.1 × 0.5–0.7 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis equisetoides

Phenology Fruiting spring–winter (Mar–Dec). Fruiting late spring–late fall.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Fresh ponds, lakes, marshes, streams, ditches, cypress swamps
Elevation 0–1500 m (0–4900 ft) 10–500 m (0–1600 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
AL; CA; CT; FL; GA; IN; LA; MA; MD; MI; MO; MS; NC; NY; RI; SC; TX; VA; WI; ON
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Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

We have not seen voucher specimens for reports of Eleocharis equistoides from Arkansas and Tennessee. It is probably extirpated from Illinois, Missouri, and Wisconsin.

Contrary to previous decriptions, sectioning of the culm reveals that the distance between the septa near the spikelet cannot be used to distinguish the two septate species, E. equisetoides and E. interstincta, which when achenes are absent are better distinguished by the septate-nodulose distal parts of the culms of E. equisetiodes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar Scirpus equisetoides, E. elliottii
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) (Elliott) Torrey: Ann. Lyceum Nat. Hist. New York 3: 296. (1836)
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