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bent spike-rush, Canada spikesedge, capitate spike-rush

Arizona spikerush

Habit Plants tufted, without creeping rhizomes. Plants annual(?), densely cespitose; rhizomes obscured among culm bases, delicate, internodes very short, scales not evident.
Culms

to 45 cm × 0.2–1 mm.

mostly ascending, terete or 4-angled, 2 cm × 0.2–0.3 mm, soft.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, greenish to whitish, translucent, membranous, apex blunt.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

ovoid, 2–3 × 1 mm, apex acute;

floral scales 7–10, 5 per mm of rachilla, bright brown or colorless, medially greenish, ovate to lanceolate, not folded lengthwise, 0.8–1.2(–1.5) × 0.6–0.8 mm, midrib obscure to slightly keeled, apex acute to acuminate.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles absent;

anthers orange-brown, 0.2–0.3 mm.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

brownish, broadly ovoid to obpyriform, much less than 2 times longer than wide, with angles plus longitudinal ridges ca. 6, rather obscure, 0.5–0.55 × 0.35 mm, apex blunt, trabeculae 15–20, distinct.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

grayish or brownish, often appressed, mostly pyramidal, 0.1 × 0.1 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis cancellata

Phenology Fruiting spring–winter (Mar–Dec). Fruiting fall (Sep).
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Wet springy places
Elevation 0–1500 m (0–4900 ft) 1700 m (5600 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
TX; Mexico (San Luis Potosí)
[BONAP county map]
Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis cancellata is very similar to E. bella. The achenes of E. cancellata are relatively broader in relation to their length, the angles and ridges obscure and fewer, and the trabeculae very distinct and faintly wavy at 35X. R. McVaugh and W. R. Anderson (1974+, vol. 13) mentioned possible intermediates with E. acicularis from Mexico. Eleocharis cancellata has been reported from the flora area from three collections: (1) Texas, Jeff Davis County, A. Brant et al. 2237, 1993, originally identified as E. bella; (2) southwestern Texas or northern Mexico (probably from near San Antonio), Parry & Palmer 912, 1879–1880; and (3) Arizona, Santa Cruz County(?), C. Wright 1937, collected in 1851. Although it was also reported from New Mexico (H. K. Svenson 1929), E. cancellata probably does not occur there (R. McVaugh and W. R. Anderson 1974+, vol. 13).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23, p. 111.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Scirpidium
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) S. Watson: Proc. Amer. Acad. Arts 18: 170. (1883)
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