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bent spike-rush, Canada spikesedge, capitate spike-rush

fewflower spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial; rhizomes 0.5–1 mm thick, scales persistent, 5–8 mm, membranous, not fibrous; resting buds absent (non-resting buds on rhizome apex ellipsoid, 10 × 2–5 mm); caudices present, hard, 1–2 mm thick.
Culms

to 45 cm × 0.2–1 mm.

markedly arched, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times as wide as thick, 5–15 cm × 0.5–1 mm, firm to hard;

culm tufts not proximally bulbous.

Leaves

distal leaf sheaths persistent, firm, distally tightly sheathing, apex acute.

distal leaf sheaths brown to reddish proximally, stramineous to reddish distally, apex sometimes reddish, membranous to papery.

Spikelets

orbicular to ovoid, 1–9 × 1–4 mm, apex rounded to acute;

proximal scale without flower, not amplexicaulous;

floral scales to 125, 11–14 per mm of rachilla, tightly appressed, dark red-brown to stramineous, ovate to elliptic, 0.8–3 × 0.6–2(–2.3) mm, membranous to cartilaginous, apex rounded to acute.

3–6 × 2–4 mm;

proximal scale empty, 2.5–4 mm, 1/2 or more as long as spikelet;

floral scales 4–9 per spikelet, 3.5–5 × 2 mm.

Flowers

perianth bristles (0–)4–8, typically 7, red-brown, rarely whitish, vestigial to much exceeding tubercle, typically equaling achene, spinules few to dense;

styles 2-fid.

perianth bristles 4–7, very unequal, some shorter than 1/2 of achene, stout and smooth, others equaling or exceeding tubercle, very slender and densely spinulose;

anthers 1–1.5 mm.

Achenes

brown ripening to black, biconvex, orbicular to obpyriform, 0.5–1.1 × 0.3–0.7 mm, apex rarely constricted proximal to tubercle, very finely reticulate at 40X.

medium brown, equilaterally trigonous to compressed trigonous or some biconvex, 1.5–2.1 × 0.9–1.2 mm, beak 0.2–0.3 × 0.3–0.4 mm.

Tubercles

stramineous to whitish, umbonate to subconic, 0.2–0.4 × 0.2–0.5 mm, apex rounded to acute.

0.25–0.5 × 0.15–0.2 mm.

2n

= 10.

Eleocharis geniculata

Eleocharis bernardina

Phenology Fruiting spring–winter (Mar–Dec). Fruiting summer.
Habitat Brackish creeks, canal banks, dune depressions, hammocks, irrigation ditches, lakeshores, lagoons, mangrove thickets, maritime mud flats, ditches, salt marshes Wet meadows in conifer forests
Elevation 0–1500 m (0–4900 ft) 2100–2700 m (6900–8900 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; IL; IN; LA; MI; MS; NE; NM; NV; OH; OK; PA; TX; ON; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
CA
Discussion

The name Eleocharis caribaea (Rottbøll) S. F. Blake is considered by most contemporary authorities to be misapplied (K. L. Wilson 1990). Eleocharis geniculata has been reported from South Carolina; I have not seen a voucher.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Although usually included in Eleocharis quinqueflora, E. bernardina is clearly more like E. suksdorfiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 101. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus geniculatus, E. capitata, E. caribaea, E. dispar Scirpus bernardinus, E. pauciflora var. bernardina, E. quinqueflora var. bernardina
Name authority (Linnaeus) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 150. (1817) (Munz & I. M. Johnston) Munz: Man. S. Calif. Bot., 68, 597. (1935)
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