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bald spike-rush, red-footed spikesedge, red-stem spikerush, redfoot spike-rush, spikesedge, éléocharide à tiges rouges

Wolf's spike-rush

Habit Plants perennial, mat-forming; rhizomes evident, long, 0.5–1.5 mm thick, soft to firm, cortex often breaking loose, longer internodes 5–30 mm, scales often fugaceous, 4–8 mm, membranous, not fibrous. Plants perennial, often forming large mats; rhizomes 0.25–0.6 mm thick, internodes 1–4 cm, scales 2 mm.
Culms

terete, often with 8–12 blunt ridges when dry, 8–80 cm × 0.3–1.4 mm, firm to soft, internally spongy.

sometimes decumbent, in same plant sides variably smooth or with 1 to few acute ridges (often nearly smooth or with 1 ridge on 1 side and several ridges on the other), greatly compressed, usually inrolled when dry, rectangular in cross section, 10–40 cm × 0.3–1.5 mm, 0.2–0.5 mm thick, firm, margins often sharply acute, margins and often 1 or more ridges minutely serrulate at 20–30X.

Leaves

distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally green to stramineous, usually inflated, often callose, membranous to papery, apex often red-brown, broadly obtuse to subacute, tooth sometimes present, to 0.1 mm.

distal leaf sheaths persistent, red proximally, colorless or stramineous or whitish distally, slightly inflated, thickly membranous, apex acute.

Spikelets

ovoid to lanceoloid or nearly cylindric, 3–18 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 15–50, 4–5 per mm of rachilla, medium brown to sometimes red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, apex rounded, in distal part lanceolate, apex entire, acute, 2–3.5 × 1.5–1.7 mm, mostly carinate.

ovoid or lanceoloid, 3–9 × 1.5–2.5 mm, apex acute;

floral scales 15–30, 6 per mm of rachilla, orange-brown or often stramineous or colorless, midrib region stramineous or greenish, ovate-lanceolate, (2.2–)2.7–3.2 × 1.5 mm, midrib prominent, apex acute.

Flowers

perianth bristles 4 or absent, light brown to stramineous, stout, usually equal, equaling achene to slightly exceeding tubercle;

stamens 3;

anthers dark yellow to stramineous, 1–1.8 mm, apiculate;

styles 2-fid.

perianth bristles absent;

anthers 1.1–1.75 mm.

Achenes

not persistent, dark yellow, stramineous, or dark brown, obovoid to obpyriform, biconvex, angles obscure, 0.9–1.6 × 0.7–1.2 mm, apex rounded, neck absent to long, smooth at 30X, or sometimes finely rugulose at 10–30X with 20 or more horizontal ridges in a vertical series.

compressed-trigonous, with angles plus longitudinal ridges ca. 9–13, prominent, obovoid, mostly 2 times longer than wide, 0.7–0.9(–1.1) × (0.4–)0.5 mm, trabeculae 30–60, rather obscure and crowded.

Tubercles

brown to whitish, pyramidal, much higher than wide to, lower than wide, 0.35–0.65 × 0.2–0.6 mm.

brownish, pyramidal, usually depressed, 0.1–0.15 × 0.2–0.25 mm.

2n

= 16, 18, 19, 20.

Eleocharis erythropoda

Eleocharis wolfii

Phenology Fruiting summer. Fruiting late spring–early summer (May–Jun).
Habitat Non-calcareous or calcareous fresh or brackish shores, marshes, meadows, fens, disturbed places Ephemeral pools in open grasslands, oak woodlands on river terraces, limestone barrens
Elevation 0–2300 m (0–7500 ft) 10–500 m (0–1600 ft)
Distribution
from FNA
AK; AR; AZ; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
[WildflowerSearch map]
from FNA
AL; AR; CO; IA; IL; IN; KS; LA; MN; MO; MS; ND; NY; OH; OK; TN; TX; WI
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis erythropoda is extremely variable. Intermediates with E. palustris variant b are common in the region of sympatry in the East, and with E. macrostachya variant b in the West, and E. erythropoda is sometimes difficult to distinguish from E. uniglumis.

Eleocharis calva Torrey is an invalid name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis wolfii is presumably extirpated from Colorado, Kansas, New York (Long Island), and Ohio. It was recently rediscovered in Wisconsin. Some literature reports (e.g. from the Great Plains) are based on misidentified specimens. I have not seen specimens to verify literature reports from Alberta, Saskatchewan, Colorado, Missouri, and Nebraska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 75. FNA vol. 23, p. 110.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Scirpidium
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara
Synonyms Scirpus glaucus Scirpus wolfii
Name authority Steudel: Syn. Pl. Glumac. 2: 76. (1855) (A. Gray) A. Gray ex Britton: in H. N. Patterson, Cat. Pl. Illinois, 46. (1876)
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