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bald spike-rush, red-footed spikesedge, red-stem spikerush, redfoot spike-rush, spikesedge, éléocharide à tiges rouges

black spikerush

Habit Plants perennial, mat-forming; rhizomes evident, long, 0.5–1.5 mm thick, soft to firm, cortex often breaking loose, longer internodes 5–30 mm, scales often fugaceous, 4–8 mm, membranous, not fibrous. Plants usually annual, tufted, not stoloniferous; rhizomes absent.
Culms

terete, often with 8–12 blunt ridges when dry, 8–80 cm × 0.3–1.4 mm, firm to soft, internally spongy.

sometimes ascending, subterete, elliptic, or polygonal, (1–)3–7.5(–9) cm × 0.2–0.5 mm, very soft.

Leaves

distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally green to stramineous, usually inflated, often callose, membranous to papery, apex often red-brown, broadly obtuse to subacute, tooth sometimes present, to 0.1 mm.

distal leaf sheaths persistent or disintegrating, colorless, stramineous or green, spotted red-brown to purple, membranous, apex narrowly acute.

Spikelets

ovoid to lanceoloid or nearly cylindric, 3–18 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 15–50, 4–5 per mm of rachilla, medium brown to sometimes red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, apex rounded, in distal part lanceolate, apex entire, acute, 2–3.5 × 1.5–1.7 mm, mostly carinate.

basal spikelets absent; never proliferous, ovoid or ellipsoid, terete, 2–4.7(–8.5) × 1–2 mm, apex acute to blunt;

proximal scale empty or with flower, persistent, clasping 1/2 of culm, similar to floral scales (but 1.1–1.7 mm, midrib region broadly green);

subproximal scale with a flower;

floral scales spiraled, 10–30(–100), 10–16 per mm of rachilla, colorless, whitish, or pale brown, mottled red-brown to purple, midribs pale brown or green, ovate or elliptic, 1–1.2 × 0.45–0.6 mm, membranous, midrib prominent, apex rounded.

Flowers

perianth bristles 4 or absent, light brown to stramineous, stout, usually equal, equaling achene to slightly exceeding tubercle;

stamens 3;

anthers dark yellow to stramineous, 1–1.8 mm, apiculate;

styles 2-fid.

perianth absent;

stamens 1;

anthers 0.2–0.3 mm;

styles 3-fid.

Achenes

not persistent, dark yellow, stramineous, or dark brown, obovoid to obpyriform, biconvex, angles obscure, 0.9–1.6 × 0.7–1.2 mm, apex rounded, neck absent to long, smooth at 30X, or sometimes finely rugulose at 10–30X with 20 or more horizontal ridges in a vertical series.

colorless (appearing green-brown from enclosed seed) or whitish, sometimes spotted red-brown, obovoid, trigonous, angles prominent, 0.5–0.6 × 0.33–0.4 mm, apex constricted proximal to tubercle, smooth.

Tubercles

brown to whitish, pyramidal, much higher than wide to, lower than wide, 0.35–0.65 × 0.2–0.6 mm.

white, gray or pale brown, pyramidal or umbonate, trigonous, 0.1–0.15 × 0.15–0.2 mm.

2n

= 16, 18, 19, 20.

Eleocharis erythropoda

Eleocharis nigrescens

Phenology Fruiting summer. Fruiting summer–winter.
Habitat Non-calcareous or calcareous fresh or brackish shores, marshes, meadows, fens, disturbed places Moist terrestrial sites such as sandy and peaty soils on pond margins, ditches, pine flatwoods
Elevation 0–2300 m (0–7500 ft) 0–30 m (0–100 ft)
Distribution
from FNA
AK; AR; AZ; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
[WildflowerSearch map]
from FNA
FL; SC; Mexico; South America; West Indies; Bermuda; Africa
[BONAP county map]
Discussion

Eleocharis erythropoda is extremely variable. Intermediates with E. palustris variant b are common in the region of sympatry in the East, and with E. macrostachya variant b in the West, and E. erythropoda is sometimes difficult to distinguish from E. uniglumis.

Eleocharis calva Torrey is an invalid name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Usually all the floral scales fall from the rachilla at maturity. The achene epidermis is usually translucent, revealing the brown color of the enclosed seed.

Clarification of delimitation of Eleocharis microlepis (Grisebach) D. A. Simpson, E. setifolia, (A. Richard) A. Raynal, and E. nigrescens in the Americas requires further investigation (D. A. Simpson 1988). Specimens from North America treated herein as E. nigrescens seem to agree well with Simpson’s illustration of E. nigrescens; the anther length of 0.2–0.3 mm recorded here is closer to that of E. microlepis. Simpson (pers. comm.) has not examined specimens of either E. nigrescens or E. microlepis from North America, and lists both E. microlepis and E. setifolia from the West Indies and only E. setifolia from South America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 75. FNA vol. 23, p. 93.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus glaucus Scirpidium nigrescens
Name authority Steudel: Syn. Pl. Glumac. 2: 76. (1855) (Nees) Steudel: Syn. Pl. Glumac. 2: 77. (1854)
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