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horsetail spike-rush, horsetail spikesedge, jointed spike-rush, jointed spike-rush (spikesedge), jointed spikesedge, knotted spike-rush

spikerush

Habit Plants perennial; rhizomes 2–4 mm thick, soft to firm, longer internodes 2–8 cm, scales persistent, 7–8 mm, membranous, tubers absent. Plants perennial; rhizomes 3–4 mm thick, soft to hard, longer internodes 3–8 cm, scales more than 1 cm, tubers absent.
Culms

terete, 50–100 cm × 5–8.4 mm, soft to firm, obscurely to clearly septate-nodulose throughout, internally hollow with complete transverse septa, closer together near the spikelet;

plants never forming filiform, flaccid culms.

obscurely 3–5-angled to terete, 40–110 cm × (2–)3.6–7.5 mm, soft, not septate-nodulose, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

Leaves

distal leaf sheaths persistent, membranous to thinly papery, apex acute to slightly acuminate.

distal leaf sheaths persistent, membranous, apex acute to long-acuminate, sometimes prolonged into a slender bladelike portion to 6 cm.

Spikelets

not proliferous, 20–45 × 4–5(–8) mm;

rachilla joints bearing prominent winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (2–)2.8–4.9 mm;

floral scales 85–160, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, obovate to oblong, (4.8–)5.2–7 × 3–4.4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to obtuse.

not proliferous, 12–43 × 3.5–6 mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale without a flower, amplexicaulous, 3.6–7.5 mm;

floral scales 30–125, 1–3 per mm of rachilla, greenish to pale brown, often with pale to dark brown submarginal band or a subapical darker spot, ovate to oblong, 4.3–5.8 × 2.5–3.3 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to acute.

Flowers

perianth bristles 3–8, stramineous to reddish, slender, vestigial to 2.5 mm, shorter than to equaling achene, smooth or obscurely retrorsely spinulose;

anthers stramineous to red-striated, 2.6–3.2 mm;

styles 2–3-fid.

perianth bristles 6–7, stramineous, margins and spinules reddish to pale brown, stout, flattened, subequal, exceeding achene, 2.5–3.2 mm, coarsely retrorsely spinulose;

anthers reddish brown, 1.5–1.9 mm;

styles 3-fid or sometimes 2-fid.

Achenes

yellowish to reddish brown, obovoid, biconvex to obscurely compressed trigonous, 1.8–2.3 × 1.4–1.9 mm, obscurely sculptured or sometimes partially smooth at 10–15X, each face with 20–40 rows of linear, transversely elongated cells, lines separating cells obscure, apex with very short neck 0.6–0.8 mm wide.

pale brown, biconvex, obpyriform, 1.7–2.3 × 1.2–1.5 mm, clearly sculptured at 10–15X, each face with 10–15 rows of transversely elongated cells, often isodiametric at achene base, apex usually constricted to neck 0.5–0.6 mm wide.

Tubercles

dark brown, lamelliform, high-deltoid, 0.6–1.1 × 0.5–0.7 mm.

stramineous, lamelliform to high-pyramidal, 0.8–1.1 × 0.7–0.9 mm.

Eleocharis equisetoides

Eleocharis obtusetrigona

Phenology Fruiting late spring–late fall. Fruiting spring–fall.
Habitat Fresh ponds, lakes, marshes, streams, ditches, cypress swamps Fresh, permanent water in ditches
Elevation 10–500 m (0–1600 ft) 0–20 m (0–100 ft)
Distribution
from FNA
AL; CA; CT; FL; GA; IN; LA; MA; MD; MI; MO; MS; NC; NY; RI; SC; TX; VA; WI; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; e Mexico; South America; Central America (Nicaragua)
[BONAP county map]
Discussion

We have not seen voucher specimens for reports of Eleocharis equistoides from Arkansas and Tennessee. It is probably extirpated from Illinois, Missouri, and Wisconsin.

Contrary to previous decriptions, sectioning of the culm reveals that the distance between the septa near the spikelet cannot be used to distinguish the two septate species, E. equisetoides and E. interstincta, which when achenes are absent are better distinguished by the septate-nodulose distal parts of the culms of E. equisetiodes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In North America Eleocharis obtusetrigona is known from only five collections, two of them originally identified as E. fistulosa (Poiret) Link [= E. acutangula (Roxburgh) Schultes], which is common in Mexico and South America. The original description of E. obtusetrigona is poor and does not clearly distinguish E. obtusangula from E. acutangula. H. K. Svenson (1939) included E. obtusetrigona in E. acutangula, which can be distinguished by its acutely trigonous culms as well as its smaller floral scales and achenes. The tubercles are often spongy as in E. quadrangulata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 118.
Parent taxa Cyperaceae > Eleocharis > subg. Limnochloa Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus equisetoides, E. elliottii Limnochloa obtusetrigona, E. fistulosa var. obtusetrigona, E. mutata var. obtusetrigona
Name authority (Elliott) Torrey: Ann. Lyceum Nat. Hist. New York 3: 296. (1836) (Lindley & Nees) Steudel: Syn. Pl. Glumac. 2: 80. (1855)
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