Eleocharis equisetoides |
Eleocharis minima |
|
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horsetail spike-rush, horsetail spikesedge, jointed spike-rush, jointed spike-rush (spikesedge), jointed spikesedge, knotted spike-rush |
small spikerush |
|
Habit | Plants perennial; rhizomes 2–4 mm thick, soft to firm, longer internodes 2–8 cm, scales persistent, 7–8 mm, membranous, tubers absent. | Plants annual, tufted, often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. |
Culms | terete, 50–100 cm × 5–8.4 mm, soft to firm, obscurely to clearly septate-nodulose throughout, internally hollow with complete transverse septa, closer together near the spikelet; plants never forming filiform, flaccid culms. |
erect, ascending or arching, quadrangular, sulcate, 1–13 cm × 0.1–0.3 mm, soft. |
Leaves | distal leaf sheaths persistent, membranous to thinly papery, apex acute to slightly acuminate. |
distal leaf sheaths persistent or disintegrating, red-brown, stramineous, green, or mottled red-brown, translucent, apex narrowly acute. |
Spikelets | not proliferous, 20–45 × 4–5(–8) mm; rachilla joints bearing prominent winglike remnants of floral scales; proximal scale empty, amplexicaulous, (2–)2.8–4.9 mm; floral scales 85–160, 1–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, obovate to oblong, (4.8–)5.2–7 × 3–4.4 mm, cartilaginous, often membranous toward margins, margins broadly translucent, membranous, apex rounded to obtuse. |
basal spikelets often present, bisexual; often proliferous, ovoid or ellipsoid, laterally compressed when young, but terete at maturity, 2–7 × 1–2.6 mm, acute; proximal scale empty, often persistent, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spirodistichous (superficially appearing ± spiraled), (4–)10–28, 6–9 per mm of rachilla, pale brown, mottled red-brown to purple, ovate or elliptic, 1.3–2.1 × 0.7–1 mm, membranous, midribs green and red-brown, prominent, apex rounded to obtuse. |
Flowers | perianth bristles 3–8, stramineous to reddish, slender, vestigial to 2.5 mm, shorter than to equaling achene, smooth or obscurely retrorsely spinulose; anthers stramineous to red-striated, 2.6–3.2 mm; styles 2–3-fid. |
perianth bristles 6, colorless, white, or stramineous, slightly shorter than to equaling achene, spinules retrorse, acute; stamens 3; anthers 0.4–0.5 mm; styles 3-fid. |
Achenes | yellowish to reddish brown, obovoid, biconvex to obscurely compressed trigonous, 1.8–2.3 × 1.4–1.9 mm, obscurely sculptured or sometimes partially smooth at 10–15X, each face with 20–40 rows of linear, transversely elongated cells, lines separating cells obscure, apex with very short neck 0.6–0.8 mm wide. |
greenish or pale brown or red-brown, often clearly spotted greenish or red-brown, ellipsoid to obovoid, trigonous (or terete?), angles prominent (keeled), 0.6–0.8 × 0.4–0.5 mm, apex constricted proximal to tubercle, smooth. |
Tubercles | dark brown, lamelliform, high-deltoid, 0.6–1.1 × 0.5–0.7 mm. |
red-brown, pyramidal, trigonous, 0.1–0.3 × 0.2–0.4 mm. |
Eleocharis equisetoides |
Eleocharis minima |
|
Phenology | Fruiting late spring–late fall. | Fruiting mid spring–fall. |
Habitat | Fresh ponds, lakes, marshes, streams, ditches, cypress swamps | Freshwater, sandy and peaty shores of lakes, muddy areas |
Elevation | 10–500 m (0–1600 ft) | 0–400 m (0–1300 ft) |
Distribution |
AL; CA; CT; FL; GA; IN; LA; MA; MD; MI; MO; MS; NC; NY; RI; SC; TX; VA; WI; ON
|
FL; MD; TX; Mexico; South America; West Indies; Central America (Costa Rica, El Salvador, Guatemala, Nicaragua, Panama) |
Discussion | We have not seen voucher specimens for reports of Eleocharis equistoides from Arkansas and Tennessee. It is probably extirpated from Illinois, Missouri, and Wisconsin. Contrary to previous decriptions, sectioning of the culm reveals that the distance between the septa near the spikelet cannot be used to distinguish the two septate species, E. equisetoides and E. interstincta, which when achenes are absent are better distinguished by the septate-nodulose distal parts of the culms of E. equisetiodes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis minima is most often confused with E. bicolor and E. baldwinii (see comments under 33. E. bicolor). Although floral scales in E. minima often superficially appear to be spirally disposed, in reality they are spirodistichous, ± decussate, while in E. baldwinii they are strictly distichous in one plane. The number of floral scales per millimeter is much greater in E. minima than in E. baldwinii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 97. |
Parent taxa | Cyperaceae > Eleocharis > subg. Limnochloa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus equisetoides, E. elliottii | Chaetocyperus jamesonii, Chaetocyperus polymorphus, Chaetocyperus viviparus, E. durandii, E. jamesonii, E. minima var. ambigua, E. oropuchensis, E. savannarum, E. subtilis, E. tenuissima, E. villaricensis, E. wrightiana, Isolepis ambigua |
Name authority | (Elliott) Torrey: Ann. Lyceum Nat. Hist. New York 3: 296. (1836) | Kunth: Enum. Pl. 2: 139. (1837) |
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