Eleocharis engelmannii
Eleocharis uniglumis
Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush
creeping spike-rush, one-glumed spikesedge, onescale spikerush, slender spike-rush, éléocharide unigume
2–40 cm × 0.5–1.5(–2) mm.
terete, often with some blunt ridges when dry, (5–)10–60 cm × 0.2–1.5 mm, firm, internally spongy.
apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.
distal leaf sheaths persistent, not splitting, proximally red, distally stramineous to green, often callose, thinly papery to thickly membranous, apex often dark red-brown, obtuse to subacute, tooth absent.
lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);
proximal scale empty, encircling ca. 2/3 of culm;
floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.
ovoid to lanceoloid, 5–10 × 2–3(–4) mm, apex acute;
proximal scale amplexicaulous, entire;
subproximal scale with flower;
floral scales often spreading in fruit, 10–20, 3–4 per mm of rachilla, brown to often red-brown, midrib regions mostly stramineous to green, broadly ovate, 3–4 × 1.8–2.5 mm, entire, apex acute to obtuse, often some carinate in distal part of spikelet.
perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;
stamens (2–)3;
anthers brown to yellow, 0.3–0.7(–1) mm;
styles 2–3-fid.
perianth bristles 0–4(–5), light brown to stramineous, stout, usually unequal, rudimentary to equaling achene;
stamens 3;
anthers dark yellow to stramineous, 1.2–2 mm;
styles 2-fid.
0.9–1.1(–1.5) × 0.7–1.1 mm.
not persistent, dark yellow or medium or dark brown, ellipsoid, obovoid, or obpyriform, biconvex, angles obscure, 1.3–1.8 × 1–1.4 mm, apex rounded, neck absent or short, smooth at 30X, or sometimes finely rugulose at 10–20X with 20 or more horizontal ridges in vertical series.
depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).
brown to whitish, pyramidal, much higher than wide to slightly depressed, sometimes spongy and with vertical rows of depressions, 0.4–0.8 × 0.3–0.8 mm.
= 10.
Eleocharis engelmannii
Eleocharis uniglumis
Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Plants treated as Eleocharis uniglumis fall within the large morphologic variation of Eurasian E. uniglumis. Two subspecies and 3 varieties were recognized for northern Europe (S.-O. Strandhede 1966), and five species that were recognized by I. D. Zinserling (1935) were placed in synonymy under E. uniglumis (S.-O. Strandhede 1966). Recognition of infraspecific taxa within North American E. uniglumis is premature. Plants commonly called E. halophila or E. uniglumis var. halophila, found mostly in Atlantic Coastal brackish habitats, have floral scales that are usually narrower and more densely placed on the rachilla than plants usually called E. uniglumis, which are found mostly in the interior; some plants are intermediate in expression of these characters. The achene and tubercle shape characters used by M. L. Fernald (1950) to distinguish E. uniglumis from E. halophila are not valid. In North America, E. uniglumis is difficult to separate from E. erythropoda and E. kamtschatica, in both of which the spikelets have only the proximal scale without a flower (empty) and the proximal scale completely amplexicaulous. Eleocharis uniglumis differs from E. erythropoda only in its broader floral scales, which are less densely placed on the rachilla; it differs from E. kamtschatica only in its smaller tubercles. It is also difficult to separate from some specimens of E. macrostachya in which the spikelets have proximal scales that are sometimes completely amplexicaulous; such plants differ from E. uniglumis only in the absence of a flower in the axil of the subproximal scale of some of the spikelets. I have not seen voucher specimens for the chromosome numbers of 2n = 27 and 28 reported by S.-O. Strandhede (1967) from Massachusetts and Nebraska, which are lower than the 2n = (44–)46(47–88) reported for Europe (S.-O. Strandhede 1965).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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- Local floras: BC, OR, WA
- Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, PNW Herbaria
- WildflowerSearch
- iNaturalist (observations)
- USDA Plants Database
- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
