Eleocharis engelmannii |
Eleocharis torticulmis |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
California twisted spikerush, twist-stem spikerush |
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Habit | Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm; culm tufts not proximally bulbous. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
6–8 × 2–3 mm; proximal scale empty, 3–4 mm, shorter than to equaling spikelet; floral scales 8–10 per spikelet, 3.5–5 × 2 mm. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
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Perianth | bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose. |
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Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm; beak 0.3–0.6 mm. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
0.25–0.6 × 0.3–0.55 mm. |
Anthers | 1.8 –3 mm. |
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2n | = 10. |
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Eleocharis engelmannii |
Eleocharis torticulmis |
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Phenology | Fruiting spring–fall. | Fruiting summer (Jun–Jul). |
Habitat | Fresh shores, marshes, disturbed places | Fens, wet meadows, vernal ponds |
Elevation | 30–2400 m (100–7900 ft) | 1100 m (3600 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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CA |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 115. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Zinserlingia |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001) |
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