Eleocharis engelmannii |
Eleocharis retroflexa |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
coastal plain spikerush |
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Habit | Plants annual, tufted, mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
erect, ascending or arching, pentagonal, sulcate, 1.5–10 cm × 0.2–0.3 mm [larger], soft. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent or disintegrating, pale brown to green, red-spotted [mostly red-brown], membranous; apex acuminate. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
basal spikelets usually present, bisexual; often proliferous, ellipsoid or obovoid, laterally compressed, 1.7–3.9 × 1.2–2 mm, apex acute; proximal scale empty or with a flower, deciduous, amplexicaulous, similar to floral scales (sometimes 2.4–2.9 mm); subproximal scale with a flower; floral scales clearly distichous, 2–6 [or more], 4–6 per mm of rachilla, pale brown [marked red-brown], ovate or elliptic, 1.8–2.5 × 0.8–1.4 mm, membranous, apex rounded to obtuse, midribs green, keeled. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 6, colorless or pale brown, shorter than achenes; spinules not evident at 45X; stamens 3; anthers (0.55–)0.7 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
stramineous (to cream), obovoid, trigonous or subterete, not compressed, angles prominent, 0.8 × 0.5–0.55 mm, apex not constricted proximal to tubercle, coarsely cancellate or honeycomb-reticulate at 10–15X. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
red-brown, pyramidal, trigonous, proximally clearly to obscurely 3-lobed, lobes decurrent on achene angles, 0.3–0.35 × 0.3–0.4 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis retroflexa |
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Phenology | Fruiting spring–fall. | Fruiting summer. |
Habitat | Fresh shores, marshes, disturbed places | Freshwater ponds, stream banks, marshes, sandy or muddy soils |
Elevation | 30–2400 m (100–7900 ft) | 0–10 m (0–0 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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AL; Mexico; Central America; South America; West Indies; Bermuda; Asia (including Indonesia); Pacific Islands; Australia |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
We have seen only one collection of Eleocharis retroflexa from the flora area (Mobile, Alabama, in 1896, US). Other populations are likely in the United States Gulf States. The broad-shouldered, strongly sculptured achenes, trilobed, decurrent tubercles, and basal spikelets are distinctive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 98. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | Scirpus retroflexus, Baeothryon retroflexum, Chaetocyperus niveus, Chaetocyperus polymorphus var. depauperatus, Chaetocyperus rugulosus, Chaetocyperus viviparus, Cyperus depauperatus, E. depauperata |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | (Poiret) Urban: Symb. Antill. 2: 165. (1900) |
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