Eleocharis engelmannii |
Eleocharis occulta |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
limestone spike-rush |
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Habit | Plants perennial, densely cespitose; rhizomes concealed by persistent dead culm bases, often ascending, short, 3–5 mm thick, hard, cortex persistent, internodes very short, scales decaying to coarse fibers, 5–12 mm, papery. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
subterete to slightly compressed, less than 2 times wider than thick, with 4–7 blunt ridges when dry, 27–56 cm × 0.2–0.5(–0.7) mm, firm to hard, spongy. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent, not splitting, proximally red, distally green to stramineous, often inflated, thinly papery to membranous, sometimes translucent, apex broadly obtuse to subtruncate, often callose, tooth absent. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid, 3–10 × 2–3 mm, apex acute; proximal scale amplexicaulous, apex 2-fid; subproximal scale with a flower; floral scales spreading in fruit, 20–50, 7 per mm of rachilla, medium brown, midrib region often narrowly stramineous, carinate, lanceolate-attenuate, 2–2.8 × 1 mm, apex 2-fid. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 3 or absent, stramineous to pale brown, rudimentary to 1/2 achene length, obscurely retrorsely spinulose; stamens 3; anthers orange-brown, 0.7–1.3 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
falling with scales, medium or dark brown, obpyriform, nearly equilaterally obscurely trigonous or cross section nearly circular, 0.7–1 × 0.5–0.65 mm, neck distinct or rarely absent, obscurely rugulose at 10–30X, 30 or more low, blunt horizontal ridges in vertical series. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
brown, depressed-pyramidal, often rudimentary, 0.1–0.15 × 0.2 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis occulta |
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Phenology | Fruiting spring–fall. | Fruiting spring (Mar–May), sometimes summer (Jul). |
Habitat | Fresh shores, marshes, disturbed places | Seasonally wet, calcareous seeps, depressions, swales, rock crevises, rocky stream beds, stream banks, wet meadows, pond margins, often on limestone |
Elevation | 30–2400 m (100–7900 ft) | 80–300 m (300–1000 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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OK; TX |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis occulta is very invariable in contrast to the extreme variability of E. compressa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 83. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | S. G. Smith: Novon 11: 247, figs. 2, 3E–K. (2001) |
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