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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush

neat spike-rush, quill spike-rush, quill spikesedge, slender spikerush, éléocharide brillante

Habit Plants perennial, mat-forming; rhizomes evident, 0.3–0.5 mm thick, hard, cortex persistent, longer internodes 2 mm, scales persistent or fugaceous, 2–3 mm, membranous to papery, slightly fibrous.
Culms

2–40 cm × 0.5–1.5(–2) mm.

terete, 2–15 cm × 0.15–0.3 mm, firm to soft.

Leaves

apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.

distal leaf sheaths persistent, not splitting, proximally stramineous to reddish, distally green or stramineous to reddish, membranous, apex often red, obtuse to acute, tooth absent.

Spikelets

lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);

proximal scale empty, encircling ca. 2/3 of culm;

floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.

ovoid, 1–4 × 1–2 mm, obtuse to acute;

proximal scale amplexicaulous, apex entire;

subproximal scale with flower;

floral scales spreading in fruit, 5–30, 8 per mm of rachilla, medium to very dark brown, midrib region often pale or greenish, broadly ovate, 1–1.3 × 1 mm, apex rounded, entire, not carinate.

Flowers

perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;

stamens (2–)3;

anthers brown to yellow, 0.3–0.7(–1) mm;

styles 2–3-fid.

perianth bristles absent;

stamens 3;

anthers yellow, 0.3 mm;

styles 3-fid.

Achenes

0.9–1.1(–1.5) × 0.7–1.1 mm.

persistent after scales fall, to dark yellow-orange or brown, broadly obpyriform, trigonous, angles evident, 0.6–0.65 × 0.5–0.55 mm, rugulose at 20–30X, 20 blunt horizontal ridges in each vertical series.

Tubercles

depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).

brown, greatly depressed, rudimentary, 0.05–15 × 0.15–0.3 mm.

2n

= 10.

Eleocharis engelmannii

Eleocharis nitida

Phenology Fruiting spring–fall. Fruiting late spring (Jun)–summer.
Habitat Fresh shores, marshes, disturbed places Fresh bog pools, streams, disturbed places
Elevation 30–2400 m (100–7900 ft) 30–400 m (100–1300 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MI; MN; NH; WI; NL; NS; ON; QC; SK
[BONAP county map]
Discussion

Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis nitida is much like E. elliptica but all structures are smaller; intermediates are unknown.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 104. FNA vol. 23, p. 86.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa
Name authority Steudel: Syn. Pl. Glumac. 2: 79. (1855) Fernald: Rhodora 8: 129. (1906)
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