Eleocharis engelmannii |
Eleocharis nitida |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
neat spike-rush, quill spike-rush, quill spikesedge, slender spikerush, éléocharide brillante |
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Habit | Plants perennial, mat-forming; rhizomes evident, 0.3–0.5 mm thick, hard, cortex persistent, longer internodes 2 mm, scales persistent or fugaceous, 2–3 mm, membranous to papery, slightly fibrous. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
terete, 2–15 cm × 0.15–0.3 mm, firm to soft. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent, not splitting, proximally stramineous to reddish, distally green or stramineous to reddish, membranous, apex often red, obtuse to acute, tooth absent. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid, 1–4 × 1–2 mm, obtuse to acute; proximal scale amplexicaulous, apex entire; subproximal scale with flower; floral scales spreading in fruit, 5–30, 8 per mm of rachilla, medium to very dark brown, midrib region often pale or greenish, broadly ovate, 1–1.3 × 1 mm, apex rounded, entire, not carinate. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles absent; stamens 3; anthers yellow, 0.3 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
persistent after scales fall, to dark yellow-orange or brown, broadly obpyriform, trigonous, angles evident, 0.6–0.65 × 0.5–0.55 mm, rugulose at 20–30X, 20 blunt horizontal ridges in each vertical series. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
brown, greatly depressed, rudimentary, 0.05–15 × 0.15–0.3 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis nitida |
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Phenology | Fruiting spring–fall. | Fruiting late spring (Jun)–summer. |
Habitat | Fresh shores, marshes, disturbed places | Fresh bog pools, streams, disturbed places |
Elevation | 30–2400 m (100–7900 ft) | 30–400 m (100–1300 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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AK; MI; MN; NH; WI; NL; NS; ON; QC; SK |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis nitida is much like E. elliptica but all structures are smaller; intermediates are unknown. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 86. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Fernald: Rhodora 8: 129. (1906) |
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