Eleocharis engelmannii |
Eleocharis nana |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
hair-like spikerush |
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Habit | Plants probably perennial, tufted; rhizomes present, ± horizontal, 0.3–0.5 mm thick, soft, longer internodes 5 mm, scales disintegrating, translucent or not. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
erect to ascending, ± quadrangular to pentagonal or broadly elliptic, sometimes sulcate, (1.9–)4.2–8.6 cm (extraterritorial plants to 12 cm) × 0.2–0.4 mm, soft. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent or fugaceous, pale brown or red-brown sometimes spotted or streaked red-brown, translucent, membranous, apex acute. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
basal spikelets absent; never proliferous, ovoid, ellipsoid or obovoid, laterally compressed when young, but terete at maturity, 1.9–4 × 1–3 mm, apex acute, proximal scale empty, deciduous, amplexicaulous or nearly so, similar to floral scales, 0.6–2.5 × 0.8–1.2 mm, midrib sometimes slightly prolonged beyond lamina; subproximal scale with a flower; floral scales distichous or spirodistichous, (2–) 4–10, 3–5 per mm of rachilla, pale brown, spotted or streaked red-brown, ovate or elliptic, 1.7–2.2 × 0.4–0.7 mm, membranous, midrib green, spotted red-brown, prominent, apex rounded. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 6, red-brown, equaling to much exceeding tubercle; spinules sparse, retrorse, sharply acute; stamens 3; anthers 0.5–0.7 × 0.1 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
whitish or pale brown, circular in outline, ellipsoid, or obovoid, trigonous, angles prominent, 0.6–0.8 × 0.5–0.6(–0.7) mm, apex constricted proximal to tubercle, smooth. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
white or red-brown, pyramidal, trigonous, 0.2–0.4 × 0.3–0.4 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis nana |
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Phenology | Fruiting spring–fall. | Fruiting summer–winter. |
Habitat | Fresh shores, marshes, disturbed places | Freshwater, moist terrestrial sites such as sandy and peaty ponds, cypress swamps |
Elevation | 30–2400 m (100–7900 ft) | 10–50 m (0–200 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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FL; South America |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. K. Svenson (1957) described Eleocharis nana as “annual (?),” but the presence of rhizomes suggests that this species may be perennial. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 95. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | Chaetocyperus punctatus, E. camptotricha, E. punctata, Scirpus camptotrichus |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Kunth: Enum. Pl. 2: 140. (1837) |
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