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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush

mountain spikerush

Habit Plants perennial, densely tufted or mat-forming; rhizomes mostly hidden by culms and roots, fairly long, 3 mm thick, hard, cortex persistent, longer internodes to 3 mm, scales persistent, ca. 8 mm, membranous, slightly fibrous.
Culms

2–40 cm × 0.5–1.5(–2) mm.

terete, when dry with few to many blunt ridges, 10–70 cm × 0.5–2 mm, soft to firm, internally hollow with complete transverse septa 2–5 mm apart, usually evident externally except in narrowest culms.

Leaves

apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.

distal leaf sheaths persistent, not splitting, proximally dark red, distally green or brown, slightly callose, papery, apex obtuse, tooth present, 0.3–1 mm.

Spikelets

lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);

proximal scale empty, encircling ca. 2/3 of culm;

floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.

ovoid, 6–21 × 3–4 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 100–500+, (15–)30–40 per mm of rachilla, medium brown to colorless, midrib regions greenish to colorless, ovate, 1.5–2.5 × 1–1.5 mm, entire, apex rounded to subacute, carinate in distal part of spikelet.

Flowers

perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;

stamens (2–)3;

anthers brown to yellow, 0.3–0.7(–1) mm;

styles 2–3-fid.

perianth bristles 6–8, pale brown, medium stout, from less than 1/2 achene length to sometimes slightly exceeding tubercle, retrorsely spinulose;

stamens 1;

anthers dark yellow to brown, 0.6–1 mm;

styles 3-fid or some 2-fid.

Achenes

0.9–1.1(–1.5) × 0.7–1.1 mm.

falling with scales, dark green or medium brown, obovoid to obpyriform, biconvex or sometimes some compressedtrigonous in same spikelet, lateral angles prominent, abaxial angle absent or evident, not prominent, 0.8–1.1 × 0.7–0.8 mm, neck short or absent, finely cancellate at 10–20X, sometimes finely rugulose, with 15 horizontal ridges in vertical series.

Tubercles

depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).

brown, pyramidal, mostly depressed, 0.2–0.35 × 0.2–0.4 mm.

2n

= 10.

Eleocharis engelmannii

Eleocharis montana

Phenology Fruiting spring–fall. Fruiting winter–fall.
Habitat Fresh shores, marshes, disturbed places Fresh temporary or artificial ponds, ditches, burned savannas, swamp margins
Elevation 30–2400 m (100–7900 ft) 20–90[–2800] m (100–300[–9200] ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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[BONAP county map]
from FNA
AL; AZ; FL; GA; LA; TX; Mexico; Central America; South America; West Indies
[BONAP county map]
Discussion

Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The taxonomy of the septate-culmed species here treated as Eleocharis montana and E. ravenelii should be evaluated. According to H. K. Svenson (1957), the type of E. montana from near Bogotá, Colombia, is the mountain extreme of the species; it has swollen culms with no visible septation.

Specimens from Acadia and St. Landry parishes, Louisiana, are intermediate between Eleocharis montana and E. montevidensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 104. FNA vol. 23, p. 79.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa Scirpus montanus, E. montana var. nodulosa, E. nodulosa
Name authority Steudel: Syn. Pl. Glumac. 2: 79. (1855) (Kunth) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 153. (1817)
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