Eleocharis engelmannii |
Eleocharis minima |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
small spikerush |
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Habit | Plants annual, tufted, often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
erect, ascending or arching, quadrangular, sulcate, 1–13 cm × 0.1–0.3 mm, soft. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent or disintegrating, red-brown, stramineous, green, or mottled red-brown, translucent, apex narrowly acute. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
basal spikelets often present, bisexual; often proliferous, ovoid or ellipsoid, laterally compressed when young, but terete at maturity, 2–7 × 1–2.6 mm, acute; proximal scale empty, often persistent, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spirodistichous (superficially appearing ± spiraled), (4–)10–28, 6–9 per mm of rachilla, pale brown, mottled red-brown to purple, ovate or elliptic, 1.3–2.1 × 0.7–1 mm, membranous, midribs green and red-brown, prominent, apex rounded to obtuse. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 6, colorless, white, or stramineous, slightly shorter than to equaling achene, spinules retrorse, acute; stamens 3; anthers 0.4–0.5 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
greenish or pale brown or red-brown, often clearly spotted greenish or red-brown, ellipsoid to obovoid, trigonous (or terete?), angles prominent (keeled), 0.6–0.8 × 0.4–0.5 mm, apex constricted proximal to tubercle, smooth. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
red-brown, pyramidal, trigonous, 0.1–0.3 × 0.2–0.4 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis minima |
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Phenology | Fruiting spring–fall. | Fruiting mid spring–fall. |
Habitat | Fresh shores, marshes, disturbed places | Freshwater, sandy and peaty shores of lakes, muddy areas |
Elevation | 30–2400 m (100–7900 ft) | 0–400 m (0–1300 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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FL; MD; TX; Mexico; South America; West Indies; Central America (Costa Rica, El Salvador, Guatemala, Nicaragua, Panama) |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis minima is most often confused with E. bicolor and E. baldwinii (see comments under 33. E. bicolor). Although floral scales in E. minima often superficially appear to be spirally disposed, in reality they are spirodistichous, ± decussate, while in E. baldwinii they are strictly distichous in one plane. The number of floral scales per millimeter is much greater in E. minima than in E. baldwinii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 97. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | Chaetocyperus jamesonii, Chaetocyperus polymorphus, Chaetocyperus viviparus, E. durandii, E. jamesonii, E. minima var. ambigua, E. oropuchensis, E. savannarum, E. subtilis, E. tenuissima, E. villaricensis, E. wrightiana, Isolepis ambigua |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Kunth: Enum. Pl. 2: 139. (1837) |
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