Eleocharis engelmannii |
Eleocharis melanocarpa |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
black-fruit spike-rush, black-fruit spikesedge |
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Culms | 2–40 cm × 0.5–1.5(–2) mm. |
to 2 times wider than thick, 35–60(–70) cm × 0.5–1.3 mm, hard, usually with to 10 blunt ribs, at 20X often finely ridged and minutely granular. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths brown to reddish. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
3–12 × 3–4 mm; proximal scale amplexicaulous, 3.5–4 mm, midrib region very broad and fleshy, apex entire; subproximal scale with a flower; floral scales 30–40, 8–10 per mm of rachilla, orange-brown to stramineous, midrib region stramineous, broadly ovate, papery or sometimes membranous, 3–4 × 2 mm, apex entire, rounded. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles present, sometimes rudimentary, brown, length variable, obscurely retrorsely spinulose; anthers brown, 1.7–2.2 mm. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
subdeltoid in outline, sometimes broadly obpyriform, equilaterally trigonous, angles prominent, 0.9–1.2 × 0.7–1.1 mm, apex truncate. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
sessile, depressed-pyramidal, often apiculate, as wide as achene, 0.3–0.5 × 0.7–1.1 mm. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis melanocarpa |
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Phenology | Fruiting spring–fall. | Fruiting summer. |
Habitat | Fresh shores, marshes, disturbed places | Fresh, oligotrophic, acid, sandy or peaty, often drying shores, ponds, ditches |
Elevation | 30–2400 m (100–7900 ft) | 10–300 m (0–1000 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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AL; FL; GA; IN; MA; MI; NC; NY; RI; SC; TX; VA
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Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis melanocarpa is reported from Rhode Island; I have not seen a voucher specimen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 89. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Melanocarpae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Torrey: Ann. Lyceum Nat. Hist. New York 3: 311. (1836) |
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