Eleocharis engelmannii
Eleocharis elliptica
Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush
elliptic spike-rush, elliptic spikesedge, slender spike-rush, éléocharide elliptique
2–40 cm × 0.5–1.5(–2) mm.
subterete to sometimes compressed, to (2–)3 times as wide as thick, often with (4–)5–10 ridges or angles, 5–90 cm × 0.3–0.8 mm, firm to soft, spongy.
apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.
distal leaf sheaths persistent, not splitting, proximally dark red, distally green to stramineous or red, thinly papery to membranous, apex usually red-brown, obtuse to subacute, often callose, tooth to 0.5 mm usually present on some culms.
lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);
proximal scale empty, encircling ca. 2/3 of culm;
floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.
ovoid, 3–8 × 2–3(–4) mm, apex obtuse (to acute);
proximal scales amplexicaulous, apex entire;
subproximal scale with a flower;
floral scales spreading in fruit, 10–30, 5–7 per mm of rachilla, medium to very dark brown, midrib region often paler, ovate, 1.7–3 × 1–1.5 mm, apex rounded to obtuse (or acute), entire or shallowly notched (emarginate) or sometimes cut to 0.5(–0.7) mm deep, apical colorless hyaline part at least as wide as long, 0.2–0.7 mm, carinate in distal part of spikelet.
perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;
stamens (2–)3;
anthers brown to yellow, 0.3–0.7(–1) mm;
styles 2–3-fid.
perianth bristles absent or rarely 1–3, pale brown, to 1/2 of achene length, sparsely retrorsely spinulose;
stamens 3;
anthers orange-brown, 0.8–1.7 mm;
styles 3-fid or some 2-fid.
0.9–1.1(–1.5) × 0.7–1.1 mm.
persistent after scales fall, usually lemon yellow or dark yellow, orange, or medium brown, rarely greenish, obpyriform, nearly equilaterally to compressed-trigonous, often some biconvex, angles evident to prominent, 0.7–1.2 × 0.6–0.9 mm, neck short or absent, finely to coarsely rugulose at 10X, 12–20 horizontal ridges in vertical series, usually finely cancellate at 10–20X.
depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).
brown to whitish, greatly depressed, apiculate, sometimes pyramidal, 0.1–0.25 × 0.25–0.45 mm.
= 10.
= 38.
Eleocharis engelmannii
Eleocharis elliptica
Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Eleocharis elliptica is very variable in rhizome thickness, internode length, culm thickness and cross section shape, presence of a tooth on the distal leaf sheath, floral scale shape, stigma number, and achene color, cross section shape, and surface sculpturing. It is difficult to separate E. elliptica from E. tenuis and E. compressa.
Three varieties have been recognized: (1) E. elliptica var. elliptica, characterized by rhizomes 0.5–1.5 mm thick with the longer internodes 2.5–10 mm, culms terete to subterete, and floral scales entire to shallowly emarginate; (2) E. elliptica var. atrata (Svenson) S. G. Smith, characterized by rhizomes 1.5–2.5 mm thick with the longer internodes very crowded to 5 mm, culms often compressed, to 2 times wider than thick, and floral scales mostly emarginate or deeply cut; 3) E. elliptica var. pseudoptera (Svenson) L. J. Harms, herein placed in E. tenuis.
Varieties of Eleocharis elliptica are not formally recognized herein because they intergrade greatly. Eleocharis elliptica var. atrata is intermediate between E. elliptica var. elliptica and E. compressa var. compressa; it has usually been placed in E. compressa but is better placed in E. elliptica because the type of var. atrata is more like E. elliptica (S. G. Smith 2001). Eleocharis tenuis var. pseudoptera Svenson might also be treated as E. elliptica var. pseudoptera following L. J. Harms (1972), who also reported that he produced fertile E. compressa × E. elliptica hybrids. I have not seen voucher specimens for any of the chromosome numbers or hybrids that Harms reported for any of the species and varieties discussed above. My field observations in calcareous fens in Wisconsin suggest that E. elliptica var. elliptica is adapted to microhabitats where the surface soil is permanently saturated by groundwater discharge; E. compressa var. compressa is adapted to microhabitats such as depressions in prairies and exposed limestone that are dry in summer; E. elliptica var. atrata is adapted to microhabitats that are intermediate between those of typical E. elliptica and those of E. compressa (S. G. Smith 2001).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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