Eleocharis engelmannii |
Eleocharis diandra |
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
Wright's spikerush, Wright's spikesedge |
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Culms | 2–40 cm × 0.5–1.5(–2) mm. |
often spreading or reclining, 2–25 cm × 0.3–1 mm. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
apex of distal leaf sheath acute to acuminate, tooth sometimes present, to 0.2 mm. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid, 2–7 × 1–4 mm, apex subacute; proximal scale either with flower or empty, base encircling 1/2 of culm; floral scales 50–100, 10 per mm of rachilla, orange to purple-brown, ovate, 1–1.5 × 0.8 mm, midrib slightly keeled, apex rounded to acute. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles absent; stamens 2(–3); anthers yellow, 0.2–0.3 mm; styles 2-fid or some 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
0.7–1 × 0.6–0.9 mm. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
deltoid 0.1–0.2 × 0.25–0.45 mm, 1/3–1/2 as high as wide, 1/8–1/4 as high and 3/4–9/10 as wide as achene. |
2n | = 10. |
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Eleocharis engelmannii |
Eleocharis diandra |
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Phenology | Fruiting spring–fall. | Fruiting late summer–fall. |
Habitat | Fresh shores, marshes, disturbed places | Fresh, mostly sandy, shores of large lakes and streams, sometimes slightly tidal |
Elevation | 30–2400 m (100–7900 ft) | 0–100 m (0–300 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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CT; MA; NH; NY; VT; ON
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Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis diandra is close to E. ovata and E. aestuum; it probably should be treated as a distinct species (A. Haines 2001). It is apparently adapted to the greatly fluctuating water levels of rivers and large lakes (e.g., Oneida Lake in New York, Lake Champlain in Vermont). I have not seen specimens of E. diandra from Maine, New Jersey, or Pennsylvania, which may be based on specimens of E. aestuum. Specimens from the Lake-of-the-Woods shore in southwest Ontario are like E. diandra; they have floral scales with apices rounded, not acute as in typical E. diandra. The only recent observations of E. diandra are from the Connecticut River in Massachusetts (1985) and Oneida Lake in New York (1968; A. Haines 2001). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 103. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | C. Wright: Bull. Torrey Bot. Club 10: 101. (1883) |
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