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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush

cylinder spikerush

Habit Plants perennial; rhizomes evident, 1 mm thick, firm, cortex unknown, internodes unknown, scales fugacious, 5 mm, membranous.
Culms

2–40 cm × 0.5–1.5(–2) mm.

terete, often with a few blunt ridges when dry, 20–50 cm × 0.2–0.7 mm, soft, spongy.

Leaves

apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.

distal leaf sheaths persistent, not splitting, proximally red or stramineous, distally green, membranous, apex usually reddish, subtruncate to obtuse, not callose, tooth present, to 0.4 mm.

Spikelets

lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);

proximal scale empty, encircling ca. 2/3 of culm;

floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.

narrowly lanceoloid, narrowly ovoid, or narrowly cylindric, 4–10 × 1–1.5 mm, apex narrowly acute;

proximal scale amplexicaulous or clasping 3/4 of culm, entire;

subproximal scale empty or with flower;

floral scales spreading in fruit, 20–80, 5–6 per mm of rachilla, orange-brown, midrib regions stramineous, narrowly ovate-lanceolate, 1.5–2 × 0.8 mm, entire, apex acute, not carinate.

Flowers

perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;

stamens (2–)3;

anthers brown to yellow, 0.3–0.7(–1) mm;

styles 2–3-fid.

perianth bristles 3–6, stramineous, stout, unequal, rudimentary to less than 1/2 achene length;

stamens 3;

anthers stramineous, 0.7–1.4 mm;

styles 3-fid.

Achenes

0.9–1.1(–1.5) × 0.7–1.1 mm.

falling with scales, stramineous, ellipsoid, compressed-trigonous, angles prominent (keeled), 0.6–0.65 × 0.4–0.45 mm, base narrowed, neck long, smooth at 40X.

Tubercles

depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).

brown, pyramidal, as high as wide to greatly depressed, 0.1–0.25 × 0.15–0.2 mm.

2n

= 10.

Eleocharis engelmannii

Eleocharis cylindrica

Phenology Fruiting spring–fall. Fruiting spring–summer.
Habitat Fresh shores, marshes, disturbed places Ephemeral pools in old rock quarry
Elevation 30–2400 m (100–7900 ft) 300 m (1000 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; South America (Argentina, Paraguay)
[BONAP county map]
Discussion

Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis cylindrica is synonymous with E. spegazzinii Barros, which was described from temperate South America (E. R. Guaglianone and O. Ueno 1990). It is very uncommon in North America, where it has been confirmed from Aransas, Burnet, and Cameron counties in Texas; reports from Lubbock and Presidio counties have not been confirmed. The report from New Mexico cannot be confirmed because the specimens lack achenes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 104. FNA vol. 23, p. 81.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa E. texana
Name authority Steudel: Syn. Pl. Glumac. 2: 79. (1855) Buckley: Proc. Acad. Nat. Sci. Philadelphia 14: 10. (1863)
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