Eleocharis engelmannii |
Eleocharis brachycarpa |
|
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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
shortfruit spikerush |
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Habit | Plants annual, densely cespitose; rhizomes obscured among culm bases, 0.2 mm thick, internodes to 4 mm, scales not evident. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
mostly ascending, 4-angled, sulcate, 5 cm × 0.1–0.2 mm, soft. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent, colorless, translucent, apex narrowly acute. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid, 2–3 × 1–2 mm, apex blunt; floral scales 10–30, ca. 10 per mm of rachilla, orange-brown to stramineous with broad colorless margin, midrib region stramineous, lanceolate, folded lengthwise, 1.5–2 × 0.8–1 mm, midrib prominent, apex acuminate. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles absent; anthers 0.7–1.2 mm. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
whitish, broadly obpyriform, much less than 2 times longer than wide, angles and longitudinal ridges ca. 6, obscure, 0.4 × 0.3 mm, apex blunt, trabeculae 20, indistinct, crowded. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
brownish, not appressed, mostly pyramidal, 0.1 × 0.1 mm. |
2n | = 10. |
|
Eleocharis engelmannii |
Eleocharis brachycarpa |
|
Phenology | Fruiting spring–fall. | Fruiting winter–spring. |
Habitat | Fresh shores, marshes, disturbed places | Wet soils |
Elevation | 30–2400 m (100–7900 ft) | 0–20 m (0–100 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
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TX; Mexico (Tamaulipas) |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Eleocharis brachycarpa is unique among the North American members of subg. Scirpidium in its floral scales, which are markedly spreading in fruit, often apparently decussate when pressed, narrowly lanceolate, and folded lengthwise so that they do not press open and flat. It is apparently very rare and local. It is admitted to the flora only on the basis of labels on a paratype (Berlandier 996) and on Berlandier 2426, both of which read “In locis paludosis – De Matamoros a las Nueces [River, Texas], Ap. 1834.” H. K. Svenson (1929) mistranscribed the locality “Nueces,” publishing it as “Mueres.” We have also seen a specimen from Mexico, Tamaulipas, 24 mi. N of San Fernando, edges of resacas, elev. 15 m, 1959, M.C. Johnston 4882C (TEX). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 111. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Scirpidium |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | Svenson: Rhodora 31: 200, plate 190, fig. 34. (1929) |
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