Eleocharis engelmannii |
Eleocharis bifida |
|
---|---|---|
Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush |
glades spikerush |
|
Habit | Plants perennial, densely cespitose; rhizomes concealed by persistent culm bases, short, 4–5 mm thick, hard, cortex persistent, internodes crowded, scales decaying to coarse fibers, 1 cm, papery. | |
Culms | 2–40 cm × 0.5–1.5(–2) mm. |
greatly compressed, 4–10 times wider than thick, often with 1 or 2 sharp ridges on 1 side, (8–)20–35 cm × 0.7–2.3 mm, hard, finely striate, spongy. |
Leaves | apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm. |
distal leaf sheaths persistent, not splitting, proximally red or stramineous, distally green to stramineous, inflated, papery, apex dark brown, broadly obtuse to subtruncate, callose, tooth absent. |
Spikelets | lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded); proximal scale empty, encircling ca. 2/3 of culm; floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute. |
ovoid, 4–9 × 2.5–4 mm, apex acute; proximal scale clasping 2/3–3/4 of culm, apex 2-fid; subproximal scale empty or with a flower; floral scales spreading in fruit, 30–60, 6–9 per mm of rachilla, medium or pale brown, midrib region often paler, ovate-lanceolate, 2.5–3.5 × 1.5 mm, apex 2-fid, carinate in distal part of spikelet. |
Flowers | perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle; stamens (2–)3; anthers brown to yellow, 0.3–0.7(–1) mm; styles 2–3-fid. |
perianth bristles 0(–5), stramineous to pale brown, 1/2 of to equaling achene length; stamens 3; anthers orange-brown, 0.7–1.5 mm; styles 3-fid. |
Achenes | 0.9–1.1(–1.5) × 0.7–1.1 mm. |
falling with scales, yellow-brown to medium brown, obovoid to obpyriform, nearly equilaterally trigonous, cross section circular to slightly compressed, angles obscure or evident, 0.9–1.1 × 0.6–0.75, neck usually very short, finely rugulose at 10–30X, 20 or more low, blunt horizontal ridges in vertical series. |
Tubercles | depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta). |
brown, depressed-pyramidal, often rudimentary, 0.1–0.25 × 0.2–0.3 mm. |
2n | = 10. |
|
Eleocharis engelmannii |
Eleocharis bifida |
|
Phenology | Fruiting spring–fall. | Fruiting spring–summer (May–Jun). |
Habitat | Fresh shores, marshes, disturbed places | Seasonally wet places on limestone, cedar (juniper) glades, stream beds, prairies, ditches |
Elevation | 30–2400 m (100–7900 ft) | 200–300 m (700–1000 ft) |
Distribution |
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
|
AL; GA; KY; TN |
Discussion | Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis bifida was previously included in E. compressa. A very few specimens of E. compressa var. compressa from the states where E. bifida is known are like E. bifida except for their entire proximal floral scales and evident rhizomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 104. | FNA vol. 23, p. 85. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa | |
Name authority | Steudel: Syn. Pl. Glumac. 2: 79. (1855) | S. G. Smith: Novon 11: 243, figs. 1F–J, 2. (2001) |
Web links |
|