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Engelman spikerush, Engelmann's spike-rush, Engelmann's spikesedge, englemann's spike-rush

Baldwin's spike-rush

Habit Plants annual(?), often mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent.
Culms

2–40 cm × 0.5–1.5(–2) mm.

often arching, sometimes spotted or streaked red-brown, quadrangular or broadly elliptic, sulcate or not, 0.5–25(–30) cm × 0.1–0.4 mm, soft.

Leaves

apex of distal leaf sheath obtuse to acute, tooth to 0.3 mm.

distal leaf sheaths persistent or disintegrating, stramineous, green, red-brown, or spotted or streaked red-brown, membranous, translucent, apex obtuse or acute.

Spikelets

lanceoloid to subcylindric or ovoid, 5–10(–20) × 2–3(–4) mm, apex acute (to rounded);

proximal scale empty, encircling ca. 2/3 of culm;

floral scales 25–100(–200), 8–12 per mm of rachilla, orangebrown to stramineous, 2(–2.5) × 1–1.3 mm, midribs mostly keeled, ovate, apex narrowly rounded to subacute.

basal spikelets usually present, pistillate; often proliferous, ovoid, ellipsoid, or linear, clearly laterally compressed, 2.3–6.9 × 0.5–2 mm, apex acute, proximal scale empty, amplexicaulous, similar to floral scales (midribs sometimes more prominent and sometimes slightly prolonged beyond lamina);

subproximal scale with a flower;

floral scales clearly distichous, 3–9, 3–4 per mm of rachilla, yellowish, pale brown, or red-brown, spotted, streaked, or mottled red-brown, narrowly ovate to oblong, (1.5–)2–5 × 0.4–0.6(–1.2) mm, membranous, apex rounded, obtuse or rarely acute, midribs keeled.

Flowers

perianth bristles present or often absent, 5–8, brown, stout, rudimentary to slightly exceeding tubercle;

stamens (2–)3;

anthers brown to yellow, 0.3–0.7(–1) mm;

styles 2–3-fid.

perianth bristles 3–7, incurved, colorless, white, or red-brown, shorter than or equaling achene;

spinules sparse, retrorse, sharply acute;

stamens 3;

anthers 1.4–2.2 mm;

styles 3-fid.

Achenes

0.9–1.1(–1.5) × 0.7–1.1 mm.

whitish or greenish, spotted dark green, obovoid, trigonous, angles prominent, 0.6–0.9 × 0.4–0.6 mm, apex constricted proximal to tubercle, smooth to finely cancellate at 10–20X.

Tubercles

depressed, subdeltoid, 0.1–0.3(–0.4) × 0.6–0.9(–1) mm, 1/10–2/5 as high as wide, 1/4 or less as high as achene, 9/10 as wide as achene, very rarely 2/5 as high as achene (in E. engelmannii var. robusta).

pale to red-brown, pyramidal, trigonous, 0.2–0.3(–0.4) × 0.2–0.5 mm.

2n

= 10.

Eleocharis engelmannii

Eleocharis baldwinii

Phenology Fruiting spring–fall. Fruiting early spring–winter.
Habitat Fresh shores, marshes, disturbed places Freshwater to moist terrestrial sites, pine savannas, cypress ponds, lakeshores
Elevation 30–2400 m (100–7900 ft) 0–60 m (0–200 ft)
Distribution
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; TN; TX; VA; WA; WI; WV; AB; BC; MB; ON
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; MS; NC; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Robust plants known only from Missouri and described as Eleocharis engelmannii var. robusta have well-developed caudices, culms to 2 mm wide (pressed), spikelets to 2 cm, floral scales to 2.5 mm and slightly cartilaginous, styles mostly trifid, achenes to 1.5 × 1 mm, and tubercles 0.3–0.5 × 0.7–1 mm and to 2/5 as high as achene. Some specimens are intermediate between E. engelmannii and E. obtusa, or rarely with E. lanceolata. I have not seen specimens to verify literature reports of E. engelmannii from Manitoba, Connecticut, Delaware, Idaho, New Hampshire, North Carolina, Oregon, Rhode Island, South Carolina, South Dakota, or Vermont.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Coarse plants of Eleocharis baldwinii usually bear more achenes per spikelet, dark colored achenes, culms gray-green when dried, and yellowish sheaths. Finer plants often bear few achenes per spikelet, whitish achenes, culms that are mid green when dried, and reddish sheaths. Achene color may relate to maturity, with darker achenes being fully mature. Although these variants appear to intergrade, they may be recognizable taxonomically. The longer floral scales in the spikelet are usually 3.5– 5 mm.

We have not seen vouchers to verify the report of Eleocharis baldwinii from Virginia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 104. FNA vol. 23, p. 98.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. engelmannii var. detonsa, E. engelmannii var. monticola, E. engelmannii var. robusta, E. monticola, E. monticola var. leviseta, E. ovata var. detonsa Chaetocyperus baldwinii
Name authority Steudel: Syn. Pl. Glumac. 2: 79. (1855) (Torrey) Chapman: Fl. South. U.S., 519. (1860)
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