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Bolander's spikerush

three-angle spikerush, three-angle spikesedge

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants perennial, mat-forming; rhizomes evident, fairly long, 1.5–3 mm thick, hard, cortex persistent, longer internodes 1.5–3 mm, scales disintegrating to fibers, 4–5 mm, papery.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

terete, often with 8–12 blunt ridges when dry, 20–55 cm × 0.3–1.3 mm, soft to hard.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths persistent, rarely splitting, proximally red to stramineous, distally green to reddish, papery, apex often dark reddish, subtruncate to obtuse, not callose, tooth present on some or all culms of plant, to 0.5 mm.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

cylindric to ovoid, 5–20 × 2–3 mm, apex rounded to subacute;

proximal scale amplexicaulous, apex entire;

subproximal scale empty;

floral scales appressed in fruit, 20–200+, 10 per mm of rachilla, dark brown, midrib regions often stramineous or greenish, oblong, 1.7–2.5 × 1 mm, apex entire, broadly rounded, usually recurved, mostly carinate.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles absent;

stamens 3;

anthers dark yellow, 1–1.5 mm;

styles 3-fid.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

falling with scales, medium to dark brown, obovoid to obpyriform, with short neck, thickly trigonous, angles prominent (keeled), 0.7–0.9 × 0.4–0.6 mm, obscurely to clearly rugulose at 10–20X with more than 20 horizontal ridges in vertical series and/or minutely cancellate at 20–30X.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

brown, rudimentary, mostly apiculate, sometimes pyramidal, 0.05–0.15 × 0.15–0.2 mm.

Eleocharis bolanderi

Eleocharis tricostata

Phenology Fruiting late spring–summer. Fruiting late spring–summer.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Wet sandy or peaty depressions, pond margins, pine barrens, savannas, mostly coastal plains
Elevation 1000–3400 m (3300–11200 ft) 10–200 m (0–700 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; MA; MI; NC; NJ; NY; SC; VA
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[BONAP county map]
Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

I have not seen specimens to verify reports of Eleocharis tricostata from Louisiana. In the absence of rhizomes and achenes, E. tricostata is easily mistaken for E. montevidensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 88.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) Torrey: Ann. Lyceum Nat. Hist. New York. 3: 310. (1836)
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