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Bolander's spikerush

California twisted spikerush, twist-stem spikerush

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm;

culm tufts not proximally bulbous.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

6–8 × 2–3 mm;

proximal scale empty, 3–4 mm, shorter than to equaling spikelet;

floral scales 8–10 per spikelet, 3.5–5 × 2 mm.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

Perianth

bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm;

beak 0.3–0.6 mm.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

0.25–0.6 × 0.3–0.55 mm.

Anthers

1.8 –3 mm.

Eleocharis bolanderi

Eleocharis torticulmis

Phenology Fruiting late spring–summer. Fruiting summer (Jun–Jul).
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Fens, wet meadows, vernal ponds
Elevation 1000–3400 m (3300–11200 ft) 1100 m (3600 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 115.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001)
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