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Bolander's spikerush

beak spike-rush, beak spikesedge, walking sedge, walking spikerush

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants densely tufted, mat-forming by means of rooting culm tips.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

1.5–3 times as wide as thick, 20–100 cm × 0.35–2 mm, firm to hard, wiry, with to 8 subacute ribs, rarely nearly smooth;

some culms arching or decumbent and rooting at tips.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths not splitting abaxially, proximally dark red to brown, apex usually reddish.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

ovoid, 5–17 × 2.5–5 mm, apex acute;

spikelets on stolons rudimentary, non-flowering, proliferous when rooting;

proximal scale empty, amplexicaulous, ovate, 2–4 mm;

subproximal scale with flower;

floral scales 20–40, 2–3 per mm of rachilla, stramineous to medium brown, midrib region paler, ovate, 3.5–6 × 2–3 mm, membranous to cartilaginous, apex entire, rounded to subacute.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles brown, equaling achene or tubercle, densely spinulose;

anthers brown, 2–2.4 mm.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

often very variable within one plant, ovoid to obovoid or obpyriform, 1.5–2.5 × 1–1.2 mm, beak to 1 × 0.6 mm.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

when present pale to dark brown, pyramidal, to 0.5 × 0.3 mm.

Eleocharis bolanderi

Eleocharis rostellata

Phenology Fruiting late spring–summer. Fruiting late spring in south, summer–fall in north.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Very wet calcareous or brackish fens, springs, shores
Elevation 1000–3400 m (3300–11200 ft) 50–2400 m (200–7900 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CT; DE; FL; ID; IL; IN; KS; LA; MA; MD; ME; MI; MN; MT; NC; NE; NJ; NM; NV; NY; OH; OK; PA; RI; SC; SD; TX; UT; VA; WA; WI; WY; BC; NS; ON; Mexico; West Indies (Cuba, Haiti, Puerto Rico)
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[BONAP county map]
Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis rostellata is highly competitive, often forming large monospecific colonies. The South American E. platypus C. B. Clarke is often treated as a synonym of E. rostellata. Eleocharis rostellata superfically closely resembles E. suksdorfiana in its culms, spikelets, and achenes, but differs in the absence of creeping rhizomes, presence of stoloniferous culms, absence of a flower in the proximal scale, and achene surface details. The collection of E. rostellata I have seen from Miami-Dade County, Florida, is from 1877. I have not seen vouchers for Archuleta County, Colorado, by H. D. Harrington (1954), or for the localities in Montana and South Carolina, which are based on the map in H. K. Svenson (1934).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 90.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Rostellatae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi Scirpus rostellatus
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) (Torrey) Torrey: Fl. New York 2: 347. (1843)
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