Eleocharis bolanderi |
Eleocharis retroflexa |
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Bolander's spikerush |
coastal plain spikerush |
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Habit | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | Plants annual, tufted, mat-forming, often stoloniferous, sometimes entirely vegetative; rhizomes absent. |
Culms | subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
erect, ascending or arching, pentagonal, sulcate, 1.5–10 cm × 0.2–0.3 mm [larger], soft. |
Leaves | distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
distal leaf sheaths persistent or disintegrating, pale brown to green, red-spotted [mostly red-brown], membranous; apex acuminate. |
Spikelets | ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
basal spikelets usually present, bisexual; often proliferous, ellipsoid or obovoid, laterally compressed, 1.7–3.9 × 1.2–2 mm, apex acute; proximal scale empty or with a flower, deciduous, amplexicaulous, similar to floral scales (sometimes 2.4–2.9 mm); subproximal scale with a flower; floral scales clearly distichous, 2–6 [or more], 4–6 per mm of rachilla, pale brown [marked red-brown], ovate or elliptic, 1.8–2.5 × 0.8–1.4 mm, membranous, apex rounded to obtuse, midribs green, keeled. |
Flowers | perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
perianth bristles 6, colorless or pale brown, shorter than achenes; spinules not evident at 45X; stamens 3; anthers (0.55–)0.7 mm; styles 3-fid. |
Achenes | falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
stramineous (to cream), obovoid, trigonous or subterete, not compressed, angles prominent, 0.8 × 0.5–0.55 mm, apex not constricted proximal to tubercle, coarsely cancellate or honeycomb-reticulate at 10–15X. |
Tubercles | whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
red-brown, pyramidal, trigonous, proximally clearly to obscurely 3-lobed, lobes decurrent on achene angles, 0.3–0.35 × 0.3–0.4 mm. |
Eleocharis bolanderi |
Eleocharis retroflexa |
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Phenology | Fruiting late spring–summer. | Fruiting summer. |
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | Freshwater ponds, stream banks, marshes, sandy or muddy soils |
Elevation | 1000–3400 m (3300–11200 ft) | 0–10 m (0–0 ft) |
Distribution |
CA; CO; ID; NV; OR; UT
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AL; Mexico; Central America; South America; West Indies; Bermuda; Asia (including Indonesia); Pacific Islands; Australia |
Discussion | Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
We have seen only one collection of Eleocharis retroflexa from the flora area (Mobile, Alabama, in 1896, US). Other populations are likely in the United States Gulf States. The broad-shouldered, strongly sculptured achenes, trilobed, decurrent tubercles, and basal spikelets are distinctive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 98. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. montevidensis var. bolanderi | Scirpus retroflexus, Baeothryon retroflexum, Chaetocyperus niveus, Chaetocyperus polymorphus var. depauperatus, Chaetocyperus rugulosus, Chaetocyperus viviparus, Cyperus depauperatus, E. depauperata |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | (Poiret) Urban: Symb. Antill. 2: 165. (1900) |
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