Eleocharis bolanderi |
Eleocharis nana |
|
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Bolander's spikerush |
hair-like spikerush |
|
Habit | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | Plants probably perennial, tufted; rhizomes present, ± horizontal, 0.3–0.5 mm thick, soft, longer internodes 5 mm, scales disintegrating, translucent or not. |
Culms | subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
erect to ascending, ± quadrangular to pentagonal or broadly elliptic, sometimes sulcate, (1.9–)4.2–8.6 cm (extraterritorial plants to 12 cm) × 0.2–0.4 mm, soft. |
Leaves | distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
distal leaf sheaths persistent or fugaceous, pale brown or red-brown sometimes spotted or streaked red-brown, translucent, membranous, apex acute. |
Spikelets | ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
basal spikelets absent; never proliferous, ovoid, ellipsoid or obovoid, laterally compressed when young, but terete at maturity, 1.9–4 × 1–3 mm, apex acute, proximal scale empty, deciduous, amplexicaulous or nearly so, similar to floral scales, 0.6–2.5 × 0.8–1.2 mm, midrib sometimes slightly prolonged beyond lamina; subproximal scale with a flower; floral scales distichous or spirodistichous, (2–) 4–10, 3–5 per mm of rachilla, pale brown, spotted or streaked red-brown, ovate or elliptic, 1.7–2.2 × 0.4–0.7 mm, membranous, midrib green, spotted red-brown, prominent, apex rounded. |
Flowers | perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
perianth bristles 6, red-brown, equaling to much exceeding tubercle; spinules sparse, retrorse, sharply acute; stamens 3; anthers 0.5–0.7 × 0.1 mm; styles 3-fid. |
Achenes | falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
whitish or pale brown, circular in outline, ellipsoid, or obovoid, trigonous, angles prominent, 0.6–0.8 × 0.5–0.6(–0.7) mm, apex constricted proximal to tubercle, smooth. |
Tubercles | whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
white or red-brown, pyramidal, trigonous, 0.2–0.4 × 0.3–0.4 mm. |
Eleocharis bolanderi |
Eleocharis nana |
|
Phenology | Fruiting late spring–summer. | Fruiting summer–winter. |
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | Freshwater, moist terrestrial sites such as sandy and peaty ponds, cypress swamps |
Elevation | 1000–3400 m (3300–11200 ft) | 10–50 m (0–200 ft) |
Distribution |
CA; CO; ID; NV; OR; UT
|
FL; South America |
Discussion | Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. K. Svenson (1957) described Eleocharis nana as “annual (?),” but the presence of rhizomes suggests that this species may be perennial. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 95. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | E. montevidensis var. bolanderi | Chaetocyperus punctatus, E. camptotricha, E. punctata, Scirpus camptotrichus |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | Kunth: Enum. Pl. 2: 140. (1837) |
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