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Bolander's spikerush

Montevideo spike rush, sand spikerush

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants perennial, mat-forming; rhizomes evident, long, 0.7–2 mm thick, firm, cortex persistent, longer internodes 1–2 cm, scales often fugaceous, 6–8 mm, membranous, not fibrous.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

terete or cross section elliptic or rectangular, with 5–10 blunt ridges when dry, 25–50 cm × 0.5–1.2 mm, firm to hard, spongy.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths persistent, not splitting, mostly proximally dark red, distally red to brown or green, thinly papery, apex usually red-brown, often callose, subtruncate to obtuse, tooth usually present on some or all culms, to 0.9 mm.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

usually ovoid or ellipsoid to subcylindric, rarely lanceoloid, 4–12 × (1.5–)2–3 mm, apex rounded to acute;

proximal scale amplexicaulous or clasping over 3/4 of culm, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 30–100, 6–10 per mm of rachilla, orange-brown, midrib regions often greenish, oblong to ovate, 1.5–2.5 × 1–1.5 mm, apex entire, broadly rounded, sometimes acute in distal part of spikelet, usually horizontally wrinkled and recurved, often carinate in distal part of spikelet.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles 5–6(–7), stramineous to medium brown, stout, often unequal, much shorter than to equaling achene, rarely all rudimentary;

stamens 3;

anthers dark yellow to stramineous, 0.8–1.5 mm;

styles 3-fid or some 2-fid.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

falling with scales, dark brown, obovoid to obpyriform, compressed trigonous, angles evident to obscure, 0.7–1 × 0.65–0.8 mm, neck absent to sometimes long, finely rugulose at 10–30X, over 20 horizontal ridges in vertical series, and/or minutely cancellate at 20–30X.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

brown to whitish, pyramidal, as high as wide or sometimes greatly depressed, (0.1–)0.25–0.3 × 0.2–0.3 mm.

2n

= 20.

Eleocharis bolanderi

Eleocharis montevidensis

Phenology Fruiting late spring–summer. Fruiting spring–fall.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Wet soil, fresh ponds, lakes, streams, springs, seeps, marshes, ditches, grasslands
Elevation 1000–3400 m (3300–11200 ft) 0–2300 m (0–7500 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
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from FNA
AL; AZ; CA; FL; KS; LA; MS; NC; NM; OK; SC; TX; Mexico; South America
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Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Eleocharis montana was long misapplied to E. montevidensis. The type of E. palmeri, recognized as a species (H. K. Svenson 1957), is indistinguishable from specimens of E. montevidensis. Eleocharis montevidensis is extremely variable; the broadly rounded, usually wrinkled and recurved floral scales are diagnostic. Achenes often fail to form. Eleocharis montevidensis is apparently closely related to E. parishii, in which the floral scales are less densely placed on the rachilla and acute to rounded, and the spikelets are narrowly lanceoloid to cylindric. In the absence of achenes and rhizomes, some specimens of E. montevidensis are easily mistaken for E. tricostata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 80.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi E. arenicola, E. montana subsp. montevidensis, E. palmeri
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) Kunth: Enum. Pl. 2: 144. (1837)
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