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Bolander's spikerush

pale spike-rush, wrinkle-sheath spike-rush, yellow spikerush, yellow spikesedge

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants with creeping rhizomes 0.5–1 mm thick.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

3–42 cm × 0.3–0.6 mm.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths often disintegrating, thinly membranous-translucent, inflated distally, often wrinkled, apex blunt.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

ellipsoid, 1.5–9 × 1–3.5 mm, apex acute to acuminate;

proximal scale without flower, not amplexicaulous;

floral scales to 65, 5–7 per mm of rachilla, loosely appressed to appressed, elliptic, 1–3 × 0.4–1.6 mm, membranous, apex acute.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles (0–)5–8, typically 7, white to stramineous, spinules dense to few;

styles 2-fid, rarely 3-fid.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

stramineous to green or dark brown, biconvex, rarely trigonous, obovoid to obpryiform, 0.4–1.1 × 0.3–0.8 mm, very finely reticulate at 40X.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

whitish to stramineous or green, 0.2–0.7 × 0.2–0.4 mm, apex acute to acuminate.

Eleocharis bolanderi

Eleocharis flavescens

Phenology Fruiting late spring–summer.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins
Elevation 1000–3400 m (3300–11200 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CT; DE; FL; GA; ID; IL; IN; LA; MA; MD; ME; MI; MN; MS; MT; NC; NH; NJ; NY; OH; PA; RI; SC; TX; UT; VA; VT; WI; WY; NB; NS; ON; QC; South America; temperate North America; West Indies
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The varieties of Eleocharis flavescens are difficult to delimit, especially in the south, and identifications of some specimens to variety are problematic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Achenes red-brown to dark brown when ripe, 0.4–0.8(–1.1) × 0.3–0.6 mm, apex rarely highly constricted proximal to tubercle; flowers with perianth bristles typically shorter than to as long as achene.
var. flavescens
1. Achenes green to golden-brown, 0.5–1.1 × 0.4–0.8 mm, often highly constricted proximal to tubercle; flowers with perianth bristles typically longer than achene.
var. olivacea
Source FNA vol. 23. FNA vol. 23, p. 100.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Subordinate taxa
E. flavescens var. flavescens, E. flavescens var. olivacea
Synonyms E. montevidensis var. bolanderi Scirpus flavescens
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) (Poiret) Urban: Symb. Antill. 4: 116. (1903)
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