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Bolander's spikerush

bald spike-rush, red-footed spikesedge, red-stem spikerush, redfoot spike-rush, spikesedge, éléocharide à tiges rouges

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants perennial, mat-forming; rhizomes evident, long, 0.5–1.5 mm thick, soft to firm, cortex often breaking loose, longer internodes 5–30 mm, scales often fugaceous, 4–8 mm, membranous, not fibrous.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

terete, often with 8–12 blunt ridges when dry, 8–80 cm × 0.3–1.4 mm, firm to soft, internally spongy.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths persistent, not splitting, proximally red (to stramineous), distally green to stramineous, usually inflated, often callose, membranous to papery, apex often red-brown, broadly obtuse to subacute, tooth sometimes present, to 0.1 mm.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

ovoid to lanceoloid or nearly cylindric, 3–18 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 15–50, 4–5 per mm of rachilla, medium brown to sometimes red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, apex rounded, in distal part lanceolate, apex entire, acute, 2–3.5 × 1.5–1.7 mm, mostly carinate.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles 4 or absent, light brown to stramineous, stout, usually equal, equaling achene to slightly exceeding tubercle;

stamens 3;

anthers dark yellow to stramineous, 1–1.8 mm, apiculate;

styles 2-fid.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

not persistent, dark yellow, stramineous, or dark brown, obovoid to obpyriform, biconvex, angles obscure, 0.9–1.6 × 0.7–1.2 mm, apex rounded, neck absent to long, smooth at 30X, or sometimes finely rugulose at 10–30X with 20 or more horizontal ridges in a vertical series.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

brown to whitish, pyramidal, much higher than wide to, lower than wide, 0.35–0.65 × 0.2–0.6 mm.

2n

= 16, 18, 19, 20.

Eleocharis bolanderi

Eleocharis erythropoda

Phenology Fruiting late spring–summer. Fruiting summer.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Non-calcareous or calcareous fresh or brackish shores, marshes, meadows, fens, disturbed places
Elevation 1000–3400 m (3300–11200 ft) 0–2300 m (0–7500 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AR; AZ; CO; CT; DE; IA; ID; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK
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Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis erythropoda is extremely variable. Intermediates with E. palustris variant b are common in the region of sympatry in the East, and with E. macrostachya variant b in the West, and E. erythropoda is sometimes difficult to distinguish from E. uniglumis.

Eleocharis calva Torrey is an invalid name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 75.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi Scirpus glaucus
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) Steudel: Syn. Pl. Glumac. 2: 76. (1855)
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