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Bolander's spikerush

Wright's spikerush, Wright's spikesedge

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

often spreading or reclining, 2–25 cm × 0.3–1 mm.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

apex of distal leaf sheath acute to acuminate, tooth sometimes present, to 0.2 mm.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

ovoid, 2–7 × 1–4 mm, apex subacute;

proximal scale either with flower or empty, base encircling 1/2 of culm;

floral scales 50–100, 10 per mm of rachilla, orange to purple-brown, ovate, 1–1.5 × 0.8 mm, midrib slightly keeled, apex rounded to acute.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles absent;

stamens 2(–3);

anthers yellow, 0.2–0.3 mm;

styles 2-fid or some 3-fid.

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

0.7–1 × 0.6–0.9 mm.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

deltoid 0.1–0.2 × 0.25–0.45 mm, 1/3–1/2 as high as wide, 1/8–1/4 as high and 3/4–9/10 as wide as achene.

Eleocharis bolanderi

Eleocharis diandra

Phenology Fruiting late spring–summer. Fruiting late summer–fall.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Fresh, mostly sandy, shores of large lakes and streams, sometimes slightly tidal
Elevation 1000–3400 m (3300–11200 ft) 0–100 m (0–300 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
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from FNA
CT; MA; NH; NY; VT; ON
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Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis diandra is close to E. ovata and E. aestuum; it probably should be treated as a distinct species (A. Haines 2001). It is apparently adapted to the greatly fluctuating water levels of rivers and large lakes (e.g., Oneida Lake in New York, Lake Champlain in Vermont). I have not seen specimens of E. diandra from Maine, New Jersey, or Pennsylvania, which may be based on specimens of E. aestuum. Specimens from the Lake-of-the-Woods shore in southwest Ontario are like E. diandra; they have floral scales with apices rounded, not acute as in typical E. diandra. The only recent observations of E. diandra are from the Connecticut River in Massachusetts (1985) and Oneida Lake in New York (1968; A. Haines 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 103.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Ovatae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) C. Wright: Bull. Torrey Bot. Club 10: 101. (1883)
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